Wannaganosuchus

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Wannaganosuchus
Temporal range: Late Paleocene, 61.6–56  Ma [1]
Wannaganosuchus.jpg
Fossil of Wannaganosuchus brachymanus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Crocodilia
Family: Alligatoridae
Subfamily: Alligatorinae
Genus: Wannaganosuchus
Erickson, 1982
Type species
Wannaganosuchus brachymanus
Erickson, 1982

Wannaganosuchus (meaning "Wannagan crocodile", in reference to the Wannagan Creek site where it was discovered) is an extinct genus of small alligatorid crocodylian. It was found in Late Paleocene-age rocks of Billings County, North Dakota, United States.

Contents

History and description

Wannaganosuchus is based on SMM  P76.28.247, a mostly complete skull and postcranial skeleton missing some vertebrae, coracoids, part of the feet, ribs, and other pieces. A few small bony scutes are also assigned to the genus, but not to the type specimen. SMM P76.28.247 was found semi-articulated in the lower part of the Bullion Creek Formation, near the base of a lignitic clay layer deposited in a marsh setting on a floodplain. Wannaganosuchus was named in 1982 by Bruce R. Erickson. The type species is W. brachymanus; the specific name means "short forefoot". [2]

The skull of SMM P76.28.247 was low, without elevated rims over the eyes, and was 159 millimetres (6.3 in) long. The snout was short and pointed compared to Cretaceous alligatorids. Its premaxillae (the bones of the tip of the snout) had five teeth each, while the maxillae (main tooth-bearing bones of the upper jaw) had thirteen teeth each, with the fourth being the largest and the last three having broad flattened crowns. The lower jaws had twenty teeth on each side, and like the upper jaws, the last five had broad crushing crowns. The forelimbs were short (hence the specific name), and the hindlimbs were long in comparison. The scutes were extensive. Most of the scutes were keeled, but did not have spikes. [2]

Erickson regarded Wannaganosuchus as a generalized early alligatorid closer to the line leading to modern alligatorids than other more specialized early alligatorids. [2] It may be the same as Allognathosuchus . [3]

Paleoecology and paleobiology

Wannaganosuchus was found in a layer with abundant plant fossils suggesting the presence of a swamp forest in the area; taxodioid logs are common. The deposit was formed under 2 to 3 metres (6.6 to 9.8 ft) of water, and grades into a shoreline deposit about 20 metres (66 ft) away. Lilies and water ferns grew along the shore and were shaded by cypress trees. [2]

SMM P76.28.247 was found in direct association with skeletons of Borealosuchus formidabilis and the long-snouted champsosaur Champsosaurus . These three taxa probably occupied different ecological niches based on size and morphology. Wannaganosuchus was a small alligatorid, only about 1.00 metre (3.28 ft) long as an adult, much smaller than its more abundant distant relative from the same quarry, Borealosuchus (roughly 4 metres (13 ft) long). Borealosuchus would have dominated the beach zone, while Champsosaurus is interpreted as a piscivore that swam near the bottom. Wannaganosuchus is thought to have lived like the modern caiman Paleosuchus , preferring seclusion. It may have floated among clumps of water plants. The structure of the arms, legs, and tail suggest that it was more aquatic than terrestrial. The mix of tooth shapes indicate it could eat a variety of foods. [2]

Phylogeny

Recent studies have consistently resolved Wannaganosuchus as a member of Alligatorinae, although its relative placement is disputed, as shown by the cladograms below. [4] [5] [6]

Cladogram from 2018 Bona et al. study: [4]

Alligatorinae

Ceratosuchus burdoshi

Allognathosuchus polyodon

Allognathosuchus wartheni

Navajosuchus mooki

Arambourgia gaudryi

Procaimanoidea kayi

Procaimanoidea utahensis

Wannaganosuchus brachymanus

Alligator prenasalis

Alligator mcgrewi

Alligator olseni

Alligator sinensis Chinese alligator

Culebrasuchus mesoamericanus

Alligator mississippiensis American alligator

Alligator mefferdi

Alligator thomsoni

Cladogram from 2019 Massonne et al. study: [5]

Alligatorinae

Alligator prenasalis

Alligator mcgrewi

Alligator olseni

Alligator sinensis Chinese alligator

Alligator thomsoni

Alligator mefferdi

Alligator mississippiensis American alligator

Procaimanoidea utahensis

Procaimanoidea kayi

Arambourgia gaudryi

Hassiacosuchus haupti

Wannaganosuchus brachymanus

Allognathosuchus wartheni

Allognathosuchus polyodon

Cladogram from 2020 Cossette & Brochu study: [6]

Alligatorinae

Ceratosuchus burdoshi

Hassiacosuchus haupti

Navajosuchus mooki

Allognathosuchus polyodon

Alligator mcgrewi

Alligator prenasalis

Wannaganosuchus brachymanus

Arambourgia gaudryi

Procaimanoidea kayi

Procaimanoidea utahensis

Alligator olseni

Alligator sinensis Chinese alligator

Alligator mississippiensis American alligator

Alligator mefferdi

Alligator thomsoni

Related Research Articles

<span class="mw-page-title-main">Alligatoridae</span> Family of crocodilians including alligators, caimans and kin

The family Alligatoridae of crocodylians includes alligators, caimans and their extinct relatives.

<i>Ceratosuchus</i> Extinct genus of reptiles

Ceratosuchus is an extinct genus of alligatorine crocodylian from latest Paleocene rocks of Colorado's Piceance Basin and earliest Eocene rocks of Wyoming's Bighorn Basin in North America, a slice of time known as the Clarkforkian North American Land Mammal Age. Like its modern relatives, Ceratosuchus was a swamp-dwelling predator. It is named for the pair of flattened, triangular bony plates that extend from the back of its head.

<i>Boverisuchus</i> Extinct genus of reptiles

Boverisuchus is an extinct genus of planocraniid crocodyliforms known from the middle Eocene of Germany and western North America. It was a relatively small crocodyliform with an estimated total length of approximately 2.2–3.6 metres (7.2–11.8 ft).

<i>Borealosuchus</i> Extinct genus of reptiles

Borealosuchus is an extinct genus of crocodyliforms that lived from the Late Cretaceous to the Eocene in North America. It was named by Christopher Brochu in 1997 for several species that had been assigned to Leidyosuchus. The species assigned to it are: B. sternbergii, the type species, from the Maastrichtian of Colorado, Montana, North Dakota, South Dakota, and Wyoming; B. acutidentatus, from the Paleocene of Saskatchewan; B. formidabilis, from the Paleocene of North Dakota; B. griffithi, from the Paleocene of Alberta; and B. wilsoni, from the Eocene of Wyoming. B. formidabilis is particularly well-known, represented by the remains of many individuals from the Wannagan Creek site in North Dakota.

Procaimanoidea is an extinct genus of alligatorid from the Eocene of North America. It was named posthumously in 1946 by Charles W. Gilmore; the type species is P. utahensis, from the Uintan of Utah. It is based on USNM 15996, a nearly complete skull and partial left hind leg. A second species, P. kayi, was named in 1941 by C.C. Mook as a species of Hassiacosuchus, for remains from the Bridgerian of Wyoming. It was reassigned to Procaimanoidea in 1967 by Wassersug and Hecht.

<i>Leidyosuchus</i> Genus of reptiles

Leidyosuchus is an extinct genus of alligatoroid from the Late Cretaceous of Alberta. It was named in 1907 by Lawrence Lambe, and the type species is L. canadensis. It is known from a number of specimens from the middle Campanian age Dinosaur Park Formation. It was a medium-sized alligatorid, with a maximum skull length greater than 40 centimeters (16 in).

<i>Hassiacosuchus</i> Extinct genus of reptiles

Hassiacosuchus is an extinct genus of small alligatorid from the early Eocene of Germany, found at the Messel pit. It was named in 1935 by K. Weitzel, and the type species is H. haupti. A second species, H. kayi, was named in 1941 by C.C. Mook for material from the Bridgerian of Wyoming, but was reassigned to Procaimanoidea in 1967 by Wassersug and Hecht. Hassiacosuchus may be the same as Allognathosuchus; Christopher Brochu has recommended continuing to use Hassiacosuchus.

<span class="mw-page-title-main">Caiman</span> Subfamily of reptiles

A caiman is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico and Central & South America from marshes and swamps to mangrove rivers and lakes. They have scaly skin and live a fairly nocturnal existence. They are relatively small-sized crocodilians with an average maximum weight of 6 to 40 kg depending on species, with the exception of the black caiman, which can grow more than 4 m (13 ft) in length and weigh in excess of 1,000 kg. The black caiman is the largest caiman species in the world and is found in the slow-moving rivers and lakes that surround the Amazon basin. The smallest species is the Cuvier's dwarf caiman, which grows to 1.2 to 1.5 m long. There are six different species of caiman found throughout the watery jungle habitats of Central and Southern America. The average length for most of the other caiman species is about 2 to 2.5 m long.

<i>Allognathosuchus</i> Extinct genus of reptiles

Allognathosuchus is an extinct genus of alligatorine crocodylian with a complicated taxonomic history. It was named in 1921.

<i>Navajosuchus</i> Extinct genus of reptiles

Navajosuchus is an extinct genus of alligatorine crocodylian. Its fossils have been found in the Paleocene-age Nacimiento Formation of the San Juan Basin, New Mexico. It was named in 1942 by Charles C. Mook, and the original type species was N. novomexicanus. N. novomexicanus was based on AMNH 5186, a partial skull collected in 1913. Later research showed that Navajosuchus novomexicanus was the same as the earlier-named Allognathosuchus mooki. However, A. mooki does not belong to the genus Allognathosuchus, and so the name of the crocodilian becomes Navajosuchus mooki. Under whichever name is used, this animal would have been a generalized predator of the Nacimiento floodplains. It was the most common Nacimiento Formation crocodilian, found in both the Puercan and Torrejonian faunal assemblages.

Arambourgia is an extinct monotypic genus of alligatorine crocodylian from Europe. It was named in 1905 as Allognathosuchus gaudryi. It was made a separate genus Arambourgia in 1940. This was synonymized with Allognathosuchus haupti in 1990, but later reassigned as its own genus once again in 2004. Arambourgia was likely to have been part of an early dispersal event of alligatorines from North America to Europe during the Eocene epoch. Arambourgia had non-serrated teeth and a deep orienirostral snout, unlike the flatter snouts of most other alligatorids.

Bottosaurus is an extinct genus of alligatorid from the Late Cretaceous-Early Paleocene of New Jersey, Texas, and possibly North Carolina and South Carolina. Two species are currently accepted, with a third requiring re-evaluation.

<i>Brachychampsa</i> Extinct genus of reptiles

Brachychampsa is an extinct genus of alligatoroid, possibly a basal caiman. Specimens have been reported from New Mexico, Colorado, Wyoming, Montana, North and South Dakota, New Jersey, and Saskatchewan, though only those from Montana, Utah, and New Mexico are based on material sufficient to justify the referral. One specimen has been reported from the Darbasa Formation of Kazakhstan, although the species status is indeterminate for the fossil. The genus first appeared during the late Campanian stage of the Late Cretaceous and became extinct during the late Maastrichtian stage of the Cretaceous. Brachychampsa is distinguished by an enlarged fifth maxillary tooth in the upper jaw.

<i>Planocrania</i> Extinct genus of reptiles

Planocrania is an extinct genus of eusuchian crocodyliforms from what is now China. Two species are currently known to belong to the genus.

<i>Stangerochampsa</i> Extinct genus of reptiles

Stangerochampsa is an extinct genus of globidontan alligatoroid, possibly an alligatorine or a stem-caiman, from the Late Cretaceous of Alberta. It is based on RTMP.86.61.1, a skull, partial lower jaws, and partial postcranial skeleton discovered in the late Campanian–early Maastrichtian-age Horseshoe Canyon Formation. Stangerochampsa was described in 1996 by Wu and colleagues. The type species is S. mccabei. The generic name honors the Stanger family, the owners of the ranch where the specimen was found, and the species name honors James Ross McCabe, who discovered, collected, and prepared it. Stangerochampsa is described as "small to medium–sized"; the type skull is 20.0 centimetres (7.9 in) long from the tip of the snout to the occipital condyle, and is 13.0 centimetres (5.1 in) wide at its greatest, while the thigh bone is 14.2 centimetres (5.6 in) long. It had heterodont dentition, with large crushing teeth at the rear of the jaws.

<span class="mw-page-title-main">Globidonta</span> Clade of reptiles

Globidonta is a clade of alligatoroids that includes alligators, caimans, and closely related extinct forms. It is defined as a stem-based clade including Alligator mississippiensis and all forms more closely related to it than to Diplocynodon. The group's fossil range extends back into the Late Cretaceous with early alligatoroids such as Albertochampsa and Brachychampsa. Extinct globidontans were particularly common in North America and Eurasia, and their modern range also includes South America.

Culebrasuchus is an extinct, monotypic genus of caiman alligatorid known from the Early to Middle Miocene (Hemingfordian) of the Panama Canal Zone of Panama. It contains a single species, Culebrasuchus mesoamericanus.

<span class="mw-page-title-main">Planocraniidae</span> Extinct family of reptiles

Planocraniidae is an extinct family of eusuchian crocodyliforms known from the Paleogene of Asia, Europe and North America. The family was coined by Li in 1976, and contains three genera, Boverisuchus, Duerosuchus and Planocrania. Planocraniids were highly specialized crocodyliforms that were adapted to living on land. They had extensive body armor, long legs, and blunt claws resembling hooves, and are sometimes informally called "hoofed crocodiles".

<span class="mw-page-title-main">Orientalosuchina</span> Extinct clade of crocodilians

Orientalosuchina is an extinct clade of alligatoroid crocodylians from South and East Asia that lived during the Paleocene and Eocene.

Jiangxisuchus is an extinct genus of crocodylian that lived during the Late Cretaceous in what is now China. It was described in 2019, and was proposed to be a basal member of Crocodyloidea. However, another concurrent 2019 study recovered Jiangxisuchus instead as a basal member of Alligatoroidea, within the newly named clade Orientalosuchina.

References

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  3. Sullivan, R.M.; Lucas, S.G.; Tsentas, C. (1988). "Navajosuchus is Allognathosuchus". Journal of Herpetology. Journal of Herpetology, Vol. 22, No. 1. 22 (1): 121–125. doi:10.2307/1564367. JSTOR   1564367.
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