Ceratosuchus Temporal range: Late Paleocene | |
---|---|
Ceratosuchus burdoshi | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Crocodilia |
Family: | Alligatoridae |
Subfamily: | Alligatorinae |
Genus: | † Ceratosuchus Schmidt, 1938 |
Type species | |
†Ceratosuchus burdoshi Schmidt, 1938 |
Ceratosuchus is an extinct genus of alligatorine crocodylian from latest Paleocene rocks of Colorado's Piceance Basin and earliest Eocene rocks of Wyoming's Bighorn Basin in North America, a slice of time known as the Clarkforkian North American Land Mammal Age. Like its modern relatives, Ceratosuchus was a swamp-dwelling predator. It is named for the pair of flattened, triangular bony plates that extend from the back of its head.
The type species is C. burdoshi, a name chosen by the Field Museum after Theodore Burdosh discovered a nearly complete skull on an expedition to Western Colorado in 1937.
"Fortunately, a knob of bone projecting from an otherwise undistinguished piece of rock had caught the eye of Mr. Burdosh, and the block had been broughtto the Museum. When the rock was chipped away, the insignificant external lump proved to belong to a fairly complete skull of a fossil crocodilian allied to the alligators; and on one posterior corner it bore a triangualr horn-like knob which proved to be identical with the mysterious separate fragments." [2]
Ceratosuchus was named in 1938 by K. P. Schmidt for a skull from Colorado. Further remains, including additional skulls, mandibles, and cervical armor, was recovered from Wyoming by University of Michigan paleontologists and described by William Bartels in 1984. The skull, of a moderately-size alligatorine, is most notable for its horns, formed by expansion of the bones (squamosals) that formed the rear corners of the skull roof. These horns were bulbous and pointed up. There were five teeth in both of the bones that made up the tip of the snout (premaxilla), fifteen in the paired maxillae that formed the sides of the upper jaw, and twenty in both dentaries of the lower jaw. The front of the lower jaw had a flattened shape, and the teeth located here pointed partially forward, with a spade-like form. The teeth had variable shapes; the first thirteen teeth in the lower jaw were pointed, while the last seven graded from a spatulate shape to a large globular shape. Aside from the horns, the skull and particularly the lower jaw of Ceratosuchus were very similar to that of its contemporary Allognathosuchus . The neck armor had blade-like keels that may have been aligned with the skull horns. [3]
Although Ceratosuchus is the only known horned alligatorine, horns are not unknown in crocodilians; similar structures are known on two other species: Voay robustus and Crocodylus rhombifer . Bartels proposed that the horns' small size and bluntness made them unlikely weapons, and their small size also made use in a threat display unlikely. Instead, he favored their use as signals for species recognition: in this case, the horns would allow Ceratosuchus and species of Allognathosuchus to tell each other apart. [3]
Ceratosuchus is known from overbank mudstone deposits. It so far has been a rare find compared to Allognathosuchus from the same rocks. Bartels reported that only 5% of Clarkforkian crocodilians collected by the University of Michigan can be certainly assigned to Ceratosuchus (although this may be partially artificial because fragmentary specimens of Ceratosuchus could be confused with Allognathosuchus). Ceratosuchus may have been ecologically restricted in comparison to Allognathosuchus. [3]
Although the blunt posterior teeth of Allognathosuchus and Ceratosuchus have been traditionally interpreted as for feeding on mollusks or turtles, Bartels noted that these crocodilians were too small to feed on large bivalves or non-juvenile turtles, and that modern crocodilians usually swallow snails whole. Instead, he proposed that skull morphology and tooth wear better fit generalized predation on a variety of invertebrates and vertebrates. [3]
The precise placement of Ceratosuchus within Alligatoroidea is disputed. Some studies have shown it as a basal member of Alligatoridae, within the stem group Alligatorinae, [4] [5] as shown in the cladogram below. [5]
Alligatoroidea |
| |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
(stem group) |
Alternatively, other studies have recovered Ceratosuchus outside of Alligatoridae and Alligatorinae, as a basal member of Alligatoroidea within the clade Globidonta, as shown in the cladogram below. [6]
Alligatoroidea |
| |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
The family Alligatoridae of crocodylians includes alligators and caimans.
Wannaganosuchus is an extinct genus of small alligatorid crocodylian. It was found in Late Paleocene-age rocks of Billings County, North Dakota, United States.
Procaimanoidea is an extinct genus of alligatorid from the Eocene of North America. It was named posthumously in 1946 by Charles W. Gilmore; the type species is P. utahensis, from the Uintan of Utah. It is based on USNM 15996, a nearly complete skull and partial left hind leg. A second species, P. kayi, was named in 1941 by C.C. Mook as a species of Hassiacosuchus, for remains from the Bridgerian of Wyoming. It was reassigned to Procaimanoidea in 1967 by Wassersug and Hecht. Procaimanoidea was a small alligatorid, and slightly heterodont, the last four teeth on each side of the jaws having blunt tips.
Leidyosuchus is an extinct genus of alligatoroid from the Late Cretaceous of Alberta. It was named in 1907 by Lawrence Lambe, and the type species is L. canadensis. It is known from a number of specimens from the middle Campanian age Dinosaur Park Formation. It was a medium-sized alligatorid, with a maximum skull length greater than 40 centimeters (16 in).
Alligatorinae is a subfamily within the family Alligatoridae that contains the alligators, and is the sister taxon to Caimaninae. Many genera in Alligatorinae are described, but only the genus Alligator is still living, with the remaining genera extinct.
Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more closely related to the alligators than the two other groups.
Allognathosuchus is an extinct genus of alligatorine crocodylian with a complicated taxonomic history. It was named in 1921.
Navajosuchus is an extinct genus of alligatorine crocodylian. Its fossils have been found in the Paleocene-age Nacimiento Formation of the San Juan Basin, New Mexico. It was named in 1942 by Charles C. Mook, and the original type species was N. novomexicanus. N. novomexicanus was based on AMNH 5186, a partial skull collected in 1913. Later research showed that Navajosuchus novomexicanus was the same as the earlier-named Allognathosuchus mooki. However, A. mooki does not belong to the genus Allognathosuchus, and so the name of the crocodilian becomes Navajosuchus mooki. Under whichever name is used, this animal would have been a generalized predator of the Nacimiento floodplains. It was the most common Nacimiento Formation crocodilian, found in both the Puercan and Torrejonian faunal assemblages.
Asiatosuchus is an extinct genus of crocodyloid crocodilians that lived in Eurasia during the Paleogene. Many Paleogene crocodilians from Europe and Asia have been attributed to Asiatosuchus since the genus was named in 1940. These species have a generalized crocodilian morphology typified by flat, triangular skulls. The feature that traditionally united these species under the genus Asiatosuchus is a broad connection or symphysis between the two halves of the lower jaw. Recent studies of the evolutionary relationships of early crocodilians along with closer examinations of the morphology of fossil specimens suggest that only the first named species of Asiatosuchus, A. grangeri from the Eocene of Mongolia, belongs in the genus. Most species are now regarded as nomina dubia or "dubious names", meaning that their type specimens lack the unique anatomical features necessary to justify their classification as distinct species. Other species such as "A." germanicus and "A." depressifrons are still considered valid species, but they do not form an evolutionary grouping with A. grangeri that would warrant them being placed together in the genus Asiatosuchus.
Acynodon is an extinct genus of eusuchian crocodylomorph from the Late Cretaceous, with fossils found throughout Southern Europe.
Arambourgia is an extinct monotypic genus of alligatorine crocodylian from Europe. It was named in 1905 and synonymized with Allognathosuchus haupti in 1990, but later reassigned as its own genus once again in 2004. Arambourgia was likely to have been part of an early dispersal event of alligatorines from North America to Europe during the Eocene epoch. Arambourgia had non-serrated teeth and a deep orienirostral snout, unlike the flatter snouts of most other alligatorids.
Bottosaurus is an extinct genus of alligatorid from the Late Cretaceous-Early Paleocene of New Jersey, Texas, and possibly North Carolina and South Carolina. Two species are currently accepted, with a third requiring re-evaluation.
Brachychampsa is an extinct genus of alligatoroid, possibly a basal caiman. Specimens have been reported from New Mexico, Colorado, Wyoming, Montana, North and South Dakota, New Jersey, and Saskatchewan, though only those from Montana, Utah, and New Mexico are based on material sufficient to justify the referral. One specimen has been reported from the Darbasa Formation of Kazakhstan, although the species status is indeterminate for the fossil. The genus first appeared during the late Campanian stage of the Late Cretaceous and became extinct during the late Maastrichtian stage of the Cretaceous. Brachychampsa is distinguished by an enlarged fifth maxillary tooth in the upper jaw.
Stangerochampsa is an extinct genus of globidontan alligatoroid, possibly an alligatorine or a stem-caiman, from the Late Cretaceous of Alberta. It is based on RTMP.86.61.1, a skull, partial lower jaws, and partial postcranial skeleton discovered in the late Campanian–early Maastrichtian-age Horseshoe Canyon Formation. Stangerochampsa was described in 1996 by Wu and colleagues. The type species is S. mccabei. The generic name honors the Stanger family, the owners of the ranch where the specimen was found, and the species name honors James Ross McCabe, who discovered, collected, and prepared it. Stangerochampsa is described as "small to medium–sized"; the type skull is 20.03 centimetres (7.89 in) long from the tip of the snout to the occipital condyle, and is 13.0 centimetres (5.1 in) wide at its greatest, while the thigh bone is 14.2 centimetres (5.6 in) long. It had heterodont dentition, with large crushing teeth at the rear of the jaws.
Krabisuchus is an extinct genus of alligatoroid crocodylian that existed in what is now Thailand during the Eocene. It was first named by paleontologists Jeremy A. Martin and Komsorn Lauprasert in 2010, and the type species is K. siamogallicus. Fossils have been found from the Krabi Basin of southern Thailand and include mostly cranial and mandibular elements as well as some postcranial remains. Krabisuchus is currently the most well known primitive alligatoroid from Asia; previously, these animals were only represented in Asia by a few fragmentary remains from China. The fossil record of alligatoroids is much more extensive in Europe and North America, where most taxa have been described.
Globidonta is a clade of alligatoroids that includes alligators, caimans, and closely related extinct forms. It is defined as a stem-based clade including Alligator mississippiensis and all forms more closely related to it than to Diplocynodon. The group's fossil range extends back into the Late Cretaceous with early alligatoroids such as Albertochampsa and Brachychampsa. Extinct globidontans were particularly common in North America and Eurasia, and their modern range also includes South America.
Culebrasuchus is an extinct, monotypic genus of caiman alligatorid known from the Early to Middle Miocene (Hemingfordian) of the Panama Canal Zone of Panama. It contains a single species, Culebrasuchus mesoamericanus.
Planocraniidae is an extinct family of eusuchian crocodyliforms known from the Paleogene of Asia, Europe and North America. The family was coined by Li in 1976, and contains three genera, Boverisuchus, Duerosuchus and Planocrania. Planocraniids were highly specialized crocodyliforms that were adapted to living on land. They had extensive body armor, long legs, and blunt claws resembling hooves, and are sometimes informally called "hoofed crocodiles".
Orientalosuchina is an extinct clade of alligatoroid crocodylians from South and East Asia that lived during the Paleocene and Eocene.
Jiangxisuchus is an extinct genus of crocodylian that lived during the Late Cretaceous in what is now China. It was described in 2019, and was proposed to be a basal member of Crocodyloidea. However, another concurrent 2019 study recovered Jiangxisuchus instead as a basal member of Alligatoroidea, within the newly named clade Orientalosuchina.