A marsh is - according to ecological definitions - a wetland that is dominated by herbaceous rather than woody plant species. [1] More in general, the word can be used for any low-lying and seasonally waterlogged terrain. In Europe and in agricultural literature low-lying meadows that require draining and embanked polderlands are also referred to as marshes or marshland.
Marshes can often be found at the edges of lakes and streams, where they form a transition between the aquatic and terrestrial ecosystems. They are often dominated by grasses, rushes or reeds. [2] If woody plants are present they tend to be low-growing shrubs, and the marsh is sometimes called a carr. This form of vegetation is what differentiates marshes from other types of wetland such as swamps, which are dominated by trees, and mires, which are wetlands that have accumulated deposits of acidic peat. [3]
Marshes provide habitats for many kinds of invertebrates, fish, amphibians, waterfowl and aquatic mammals. [4] This biological productivity means that marshes contain 0.1% of global sequestered terrestrial carbon. [5] Moreover, they have an outsized influence on climate resilience of coastal areas and waterways, absorbing high tides and other water changes due to extreme weather. [5] Though some marshes are expected to migrate upland, most natural marshlands will be threatened by sea level rise and associated erosion. [5]
Marshes provide a habitat for many species of plants, animals, and insects that have adapted to living in flooded conditions or other environments. [1] The plants must be able to survive in wet mud with low oxygen levels. Many of these plants, therefore, have aerenchyma, channels within the stem that allow air to move from the leaves into the rooting zone. [1] Marsh plants also tend to have rhizomes for underground storage and reproduction. Common examples include cattails, sedges, papyrus and sawgrass. Aquatic animals, from fish to salamanders, are generally able to live with a low amount of oxygen in the water. Some can obtain oxygen from the air instead, while others can live indefinitely in conditions of low oxygen. [3] The pH in marshes tends to be neutral to alkaline, as opposed to bogs, where peat accumulates under more acid conditions.
Marshes provide habitats for many kinds of invertebrates, fish, amphibians, waterfowl and aquatic mammals. [4] Marshes have extremely high levels of biological production, some of the highest in the world, and therefore are important in supporting fisheries. [1]
Marshes also improve water quality by acting as a sink to filter pollutants and sediment from the water that flows through them. Marshes partake in water purification by providing nutrient and pollution consumption. [6] Marshes (and other wetlands) are able to absorb water during periods of heavy rainfall and slowly release it into waterways and therefore reduce the magnitude of flooding. [7] Marshes also provide the services of tourism, recreation, education, and research. [6]
Marshes differ depending mainly on their location and salinity. Both of these factors greatly influence the range and scope of animal and plant life that can survive and reproduce in these environments. The three main types of marsh are salt marshes, freshwater tidal marshes, and freshwater marshes. [3] These three can be found worldwide and each contains a different set of organisms.
Saltwater marshes are found around the world in mid to high latitudes, wherever there are sections of protected coastline. They are located close enough to the shoreline that the motion of the tides affects them, and, sporadically, they are covered with water. They flourish where the rate of sediment buildup is greater than the rate at which the land level is sinking. [3] Salt marshes are dominated by specially adapted rooted vegetation, primarily salt-tolerant grasses. [8]
Salt marshes are most commonly found in lagoons, estuaries, and on the sheltered side of a shingle or sandspit. The currents there carry the fine particles around to the quiet side of the spit, and sediment begins to build up. These locations allow the marshes to absorb the excess nutrients from the water running through them before they reach the oceans and estuaries. [3] These marshes are slowly declining. Coastal development and urban sprawl has caused significant loss of these essential habitats. [9]
Although considered a freshwater marsh, this form of marsh is affected by the ocean tides. However, without the stresses of salinity at work in its saltwater counterpart, the diversity of the plants and animals that live in and use freshwater tidal marshes is much higher than in salt marshes. The most serious threats to this form of marsh are the increasing size and pollution of the cities surrounding them. [3]
Ranging greatly in both size and geographic location, freshwater marshes make up the most common form of wetland in North America. They are also the most diverse of the three types of marsh. Some examples of freshwater marsh types in North America are:
Wet meadows occur in areas such as shallow lake basins, low-lying depressions, and the land between shallow marshes and upland areas. They also occur on the edges of large lakes and rivers. Wet meadows often have very high plant diversity and high densities of buried seeds. [8] [10] They are regularly flooded, but are often dry in the summer.
Vernal pools are a type of marsh found only seasonally in shallow depressions in the land. They can be covered in shallow water, but in the summer and fall, they can be completely dry. In western North America, vernal pools tend to form in open grasslands, [11] whereas in the east they often occur in forested landscapes. [12] Further south, vernal pools form in pine savannas and flatwoods. Many amphibian species depend upon vernal pools for spring breeding; these ponds provide habitat free from fish, which eat the eggs and young of amphibians. [8] An example is the endangered gopher frog. [13] Similar temporary ponds occur in other world ecosystems, where they may have local names. However, the term vernal pool can be applied to all such temporary pool ecosystems. [8]
Playa lakes are a form of shallow freshwater marsh that occurs in the southern high plains of the United States. [14] Like vernal pools, they are only present at certain times of the year and generally have a circular shape. [15] As the playa dries during the summer, conspicuous plant zonation develops along the shoreline. [16]
Prairie potholes are found in the northern parts of North America as the Prairie Pothole Region. These landscapes were once covered by glaciers, and as a result shallow depressions were formed in great numbers. These depressions fill with water in the spring. They provide important breeding habitats for many species of waterfowl. Some pools only occur seasonally, while others retain enough water to be present all year. [17]
Many kinds of marsh occur along the fringes of large rivers. The different types are produced by factors such as water level, nutrients, ice scour, and waves. [18]
Large tracts of tidal marsh have been embanked and artificially drained. They are usually known by the Dutch name of polders. In Northern Germany and Scandinavia they are called Marschland , Marsch or marsk; in France marais maritime. In the Netherlands and Belgium, they are designated as marine clay districts. In East Anglia, a region in the East of England, the embanked marshes are also known as Fens.
Some areas of the world have already lost 90% of their wetlands, including marshes. They have been drained to create agricultural land or filled to accommodate urban sprawl. Restoration is the process of returning marshes to the landscape to replace those lost in the past. [1] Restoration can be done on a large scale, such as by allowing rivers to flood naturally in the spring, or on a small scale by returning wetlands to urban landscapes.
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environments and are an example of an ecotone. Estuaries are subject both to marine influences such as tides, waves, and the influx of saline water, and to fluvial influences such as flows of freshwater and sediment. The mixing of seawater and freshwater provides high levels of nutrients both in the water column and in sediment, making estuaries among the most productive natural habitats in the world.
A swamp is a forested wetland. Swamps are considered to be transition zones because both land and water play a role in creating this environment. Swamps vary in size and are located all around the world. The water of a swamp may be fresh water, brackish water, or seawater. Freshwater swamps form along large rivers or lakes where they are critically dependent upon rainwater and seasonal flooding to maintain natural water level fluctuations. Saltwater swamps are found along tropical and subtropical coastlines. Some swamps have hammocks, or dry-land protrusions, covered by aquatic vegetation, or vegetation that tolerates periodic inundation or soil saturation. The two main types of swamp are "true" or swamp forests and "transitional" or shrub swamps. In the boreal regions of Canada, the word swamp is colloquially used for what is more formally termed a bog, fen, or muskeg. Some of the world's largest swamps are found along major rivers such as the Amazon, the Mississippi, and the Congo.
Wetlands, or simply a wetland, is a distinct ecosystem that is flooded or saturated by water, either permanently or seasonally. Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The primary factor that distinguishes wetlands from terrestrial land forms or water bodies is the characteristic vegetation of aquatic plants, adapted to the unique anoxic hydric soils. Wetlands are considered among the most biologically diverse of all ecosystems, serving as home to a wide range of plant and animal species. Methods for assessing wetland functions, wetland ecological health, and general wetland condition have been developed for many regions of the world. These methods have contributed to wetland conservation partly by raising public awareness of the functions some wetlands provide. Constructed wetlands are designed and built to treat municipal and industrial wastewater as well as to divert stormwater runoff. Constructed wetlands may also play a role in water-sensitive urban design.
A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials – often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and muskeg; alkaline mires are called fens. A baygall is another type of bog found in the forest of the Gulf Coast states in the United States. They are often covered in heath or heather shrubs rooted in the sphagnum moss and peat. The gradual accumulation of decayed plant material in a bog functions as a carbon sink.
A salt marsh, saltmarsh or salting, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. It is dominated by dense stands of salt-tolerant plants such as herbs, grasses, or low shrubs. These plants are terrestrial in origin and are essential to the stability of the salt marsh in trapping and binding sediments. Salt marshes play a large role in the aquatic food web and the delivery of nutrients to coastal waters. They also support terrestrial animals and provide coastal protection.
The littoral zone, also called litoral or nearshore, is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark, to coastal areas that are permanently submerged — known as the foreshore — and the terms are often used interchangeably. However, the geographical meaning of littoral zone extends well beyond the intertidal zone to include all neritic waters within the bounds of continental shelves.
Freshwater ecosystems are a subset of Earth's aquatic ecosystems. They include lakes, ponds, rivers, streams, springs, bogs, and wetlands. They can be contrasted with marine ecosystems, which have a larger salt content. Freshwater habitats can be classified by different factors, including temperature, light penetration, nutrients, and vegetation. There are three basic types of freshwater ecosystems: Lentic, lotic and wetlands. Freshwater ecosystems contain 41% of the world's known fish species.
A Ramsar site is a wetland site designated to be of international importance under the Ramsar Convention, also known as "The Convention on Wetlands", an international environmental treaty signed on 2 February 1971 in Ramsar, Iran, under the auspices of UNESCO. It came into force on 21 December 1975, when it was ratified by a sufficient number of nations. It provides for national action and international cooperation regarding the conservation of wetlands, and wise sustainable use of their resources. Ramsar identifies wetlands of international importance, especially those providing waterfowl habitat.
Vernal pools, also called vernal ponds or ephemeral pools, are seasonal pools of water that provide habitat for distinctive plants and animals. They are considered to be a distinctive type of wetland usually devoid of fish, and thus allow the safe development of natal amphibian and insect species unable to withstand competition or predation by fish. Certain tropical fish lineages have however adapted to this habitat specifically.
A tidal marsh is a marsh found along rivers, coasts and estuaries which floods and drains by the tidal movement of the adjacent estuary, sea or ocean. Tidal marshes experience many overlapping persistent cycles, including diurnal and semi-diurnal tides, day-night temperature fluctuations, spring-neap tides, seasonal vegetation growth and decay, upland runoff, decadal climate variations, and centennial to millennial trends in sea level and climate.
An aquatic ecosystem is an ecosystem found in and around a body of water, in contrast to land-based terrestrial ecosystems. Aquatic ecosystems contain communities of organisms—aquatic life—that are dependent on each other and on their environment. The two main types of aquatic ecosystems are marine ecosystems and freshwater ecosystems. Freshwater ecosystems may be lentic ; lotic ; and wetlands.
A wet meadow is a type of wetland with soils that are saturated for part or all of the growing season which prevents the growth of trees and brush. Debate exists whether a wet meadow is a type of marsh or a completely separate type of wetland. Wet prairies and wet savannas are hydrologically similar.
Lobelia dortmanna, Dortmann's cardinalflower or water lobelia, is a species of flowering plant in the bellflower family Campanulaceae. This stoloniferous herbaceous perennial aquatic plant with basal leaf-rosettes and flower stalks grows to 0.7–2 m (2.3–6.6 ft) tall. The flowers are 1–2 cm long, with a five-lobed white to pale pink or pale blue corolla, produced in groups of one to ten on an erect raceme held above the water surface. The fruit is a capsule 5–10 mm long and 3–5 mm wide, containing numerous small seeds.
A pond is a small, still, land-based body of water formed by pooling inside a depression, either naturally or artificially. A pond is smaller than a lake and there are no official criteria distinguishing the two, although defining a pond to be less than 5 hectares in area, less than 5 metres (16 ft) in depth and with less than 30% with emergent vegetation helps in distinguishing the ecology of ponds from those of lakes and wetlands. Ponds can be created by a wide variety of natural processes, or they can simply be isolated depressions filled by runoff, groundwater, or precipitation, or all three of these. They can be further divided into four zones: vegetation zone, open water, bottom mud and surface film. The size and depth of ponds often varies greatly with the time of year; many ponds are produced by spring flooding from rivers. Ponds may be freshwater or brackish in nature. 'Ponds' consisting of saltwater, with a direct connection to the sea to maintain full salinity, are normally regarded as part of the marine environment. These bodies of water do not support fresh or brackish water-based organisms, and are not considered to be ponds.
Freshwater biology is the scientific biological study of freshwater ecosystems and is a branch of limnology. This field seeks to understand the relationships between living organisms in their physical environment. These physical environments may include rivers, lakes, streams, ponds, lakes, reservoirs, or wetlands. Knowledge from this discipline is also widely used in industrial processes to make use of biological processes involved with sewage treatment and water purification. Water presence and flow is an essential aspect to species distribution and influences when and where species interact in freshwater environments.
Brackish marshes develop from salt marshes where a significant freshwater influx dilutes the seawater to brackish levels of salinity. This commonly happens upstream from salt marshes by estuaries of coastal rivers or near the mouths of coastal rivers with heavy freshwater discharges in the conditions of low tidal ranges.
Classification of wetlands has been a problematical task, with the commonly accepted definition of what constitutes a wetland being among the major difficulties. A number of national wetland classifications exist. In the 1970s, the Ramsar Convention on Wetlands of International Importance introduced a first attempt to establish an internationally acceptable wetland classification scheme.
A freshwater marsh is a non-tidal, non-forested marsh wetland that contains fresh water, and is continuously or frequently flooded. Freshwater marshes primarily consist of sedges, grasses, and emergent plants. Freshwater marshes are usually found near the mouths of rivers, along lakes, and are present in areas with low drainage like abandoned oxbow lakes. It is the counterpart to the salt marsh, an upper coastal intertidal zone of bio-habitat, which is regularly flushed with sea water.
Salt pannes and pools are water retaining depressions located within salt and brackish marshes. Pools tend to retain water during the summer months between high tides, whereas pannes generally do not. Salt pannes generally start when a mat of organic debris is deposited upon existing vegetation, killing it. This creates a slight depression in the surrounding vegetation which retains water for varying periods of time. Upon successive cycles of inundation and evaporation the panne develops an increased salinity greater than that of the larger body of water. This increased salinity dictates the type of flora and fauna able to grow within the panne. Salt pools are also secondary formations, though the exact mechanism(s) of formation are not well understood; some have predicted they will increase in size and abundance in the future due to rising sea levels.
Panicum hemitomon is a species of grass known by the common name maidencane. It is native to North America, where it occurs along the southeastern coastline from New Jersey to Texas. It is also present in South America.