Ancyloceratina

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Ancyloceratina
Temporal range: 199.6–64.8  Ma
Hamites.jpg
A reconstruction of a heteromorph ammonite, Hamites sp.
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Ammonoidea
Order: Ammonitida
Suborder: Ancyloceratina
Wiedmann  [ de ], 1966
Superfamilies

The Ancyloceratina were a diverse suborder of ammonite most closely related to the ammonites of order Lytoceratina. They evolved during the Late Jurassic but were not very common until the Cretaceous period, when they rapidly diversified and became one of the most distinctive components of Cretaceous marine faunas. They have been recorded from every continent and many are used as zonal or index fossils. The most distinctive feature of the majority of the Ancyloceratina is the tendency for most of them to have shells that are not regular spirals like most other ammonites. These irregularly-coiled ammonites are called heteromorph ammonites, in contrast to regularly coiled ammonites, which are called homomorph ammonites.

Contents

Biology

The biology of the heteromorph ammonites is not clear, but one certainty is that their uncoiled shells would have made these forms very poor swimmers. Open shells, particularly ones with spines and ribs, create a lot of drag; but more importantly, the orientation of the shell, with the body hanging below the buoyant part of the shell, would have created a serious impediment to efficient swimming. It's more likely these ammonites either drifted in the plankton, collecting small animals on long tentacles like modern jellyfish, or else they crawled along the sea floor feeding on sessile or slow-moving animals such as clams.

Morphology

Assemblage of Barremian heteromorph ammonites from southeastern France Ammonite cemetery.jpg
Assemblage of Barremian heteromorph ammonites from southeastern France

Ancyloceratida varied widely in size, ranging from diminutive Ptychoceras, which was barely 3 cm (1.2 in) long, to Baculites and Diplomoceras , which could grow to 1–2 m (3.3–6.6 ft) in length. Some species were very widely distributed, for example some species of Hamites can be found in Eurasia, South America, Australia, and Antarctica. Others, like those inhabiting the Western Interior Seaway that covered much of the US, were much more localized.

In the more primitive forms, the shell departs only slightly from being a perfect spiral, with only the last, outermost whorl being open, forming a hook underneath the main spiral. In such forms the spiral was the chambered, buoyant part of the shell, and the hook was the living chamber in which the soft body of the ammonite resided. Examples of such types were Ancyloceras , Protanisoceras , and Tropaeum . The more advanced heteromorphs departed from such forms radically. The shells of Ptychoceras consisted of three or four shafts squashed together and connected with tight, 180 degree bends. Members of the genus Hamites were much larger but had a similar sort of shape, though the shafts were open so that the whole thing looked rather like a big paper clip.

Many of the earlier heteromorph forms had regularly coiled shells barely distinguishable from the homomorph ammonites (for example, the Lower Cretaceous genus Deshayesites ). Some offshoots of the uncoiled varieties even went back to being regularly coiled. Most notable among these were the Scaphites , Hoploscaphites and their relatives, which were mostly regular spirals in shape except for a very slight hook at the end.

The Late Cretaceous enjoyed the widest variety of heteromorphs, including the straight-shelled Baculites and Sciponoceras ; the helically coiled Turrilites ; and the bizarre Nipponites , which looked more like a ball of string than anything else. Some forms combined different coiling modes. For example, Nostoceras started off with a helix like a Turrilites, but had a planar hook hanging underneath, more like an Ancyloceras.

The thin shelled heteromorphic ammonites probably lived at depths 36–183 m (118.1–600.4 ft). [1]

Classification

Crioceratites nolani with spines partially restored Crioceratites nolani France 450 mm.jpg
Crioceratites nolani with spines partially restored
Nipponites mirabilis Nipponites mirabilis Musee des Confluences 18 10 2015 1.jpg
Nipponites mirabilis
Scaphites sp. Scaphite 180308.jpg
Scaphites sp.

Suborder Ancyloceratina

Footnotes

  1. "Paleoecologic Context," Pasche and May (2001); page 224.

Related Research Articles

Ammonoidea Extinct subclass of cephalopod molluscs

Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids than they are to shelled nautiloids such as the living Nautilus species. The earliest ammonites appeared during the Devonian, with the last species vanishing shortly after the Cretaceous–Paleogene extinction event, during the Danian epoch of the Paleocene.

<i>Hamites</i> (genus) Genus of molluscs (fossil)

Hamites is a genus of heteromorph ammonite that evolved late in the Aptian stage of the Early Cretaceous and lasted into the Cenomanian stage of the Late Cretaceous. The genus is almost certainly paraphyletic but remains in wide use as a "catch all" for heteromorph ammonites of the superfamily Turrilitoidea that do not neatly fit into the more derived groupings. In an attempt to identify clades within the genus, it has been divided up into a series of new genera or subgenera by different palaeontologists, including Eohamites, Hamitella, Helicohamites, Lytohamites, Planohamites, Psilohamites, and Sziveshamites.

<i>Baculites</i> Genus of molluscs (fossil)

Baculites is an extinct genus of cephalopods with a nearly straight shell, included in the heteromorph ammonites. The genus, which lived worldwide throughout most of the Late Cretaceous, and which briefly survived the K-Pg mass extinction event, was named by Lamarck in 1799.

<i>Scaphites</i> Genus of mollusks (fossil)

Scaphites is a genus of heteromorph ammonites belonging to the Scaphitidae family. They were a widespread genus that thrived during the Late Cretaceous period.

<i>Allocrioceras</i> Genus of molluscs (fossil)

Allocrioceras is an ammonoid cephalopod from the Turonian to Santonian stages of the Late Cretaceous, included in the turrilitoid family Anisoceratidae. Its shell is strongly ribbed and is in the form of a widely open spiral.

<i>Ancyloceras</i> Genus of molluscs (fossil)

Ancyloceras is an extinct genus of heteromorph ammonites found throughout the world during the Lower Cretaceous, from the Lower Barremian epoch until the genus extinction during the Lower Aptian.

Anisoceratidae is an extinct family of heteromorph ammonites which belong to the Ancyloceratina superfamily Turrilitoidea. Members of the family range is from the lower Albian to the upper Turonian. The family is possibly derived from a member of the Hamitidae.

Baculitidae Extinct family of molluscs

Baculitidae is a family of extinct ammonoid cephalopods that lived mostly during the Late Cretaceous, and often included in the suborder Ancyloceratina.

Lytoceratina Extinct suborder of ammonites

Lytoceratina is a suborder of Jurassic and Cretaceous ammonites that produced loosely coiled, evolute and gyroconic shells in which the sutural element are said to have complex moss-like endings.

Turrilitidae Extinct family of ammonites

Turrilitidae is a family of extinct heteromorph ammonite cephalopods. All members had shells that coiled helically that tended to resemble auger shells. The ecological roles turrilitids played is largely unknown, as experts are still speculating what niches they filled. Some are suspected of floating in the water column, while others, such as the eponymous Turrilites, are believed to have been bottom-dwellers. The name of the type genus Turrilites is a hybrid formation based on Latin turris "tower" and Greek lithos "stone", coined by Lamarck in 1801.

Turrilitoidea Extinct superfamily of ammonites

Turrilitoidea is a diverse superfamily of Cretaceous ammonites generally considered as heteromorphic and commonly included in the suborder Ancyloceratina. Shells of this diverse group do not coil planospirally, as typical for most ammonites, but rather take on a variety of unique forms. Separation of Turrilitoidea along with Ptychoceratoidea and Scaphatoidea into a different suborder, Turrilitina, was proposed by Beznosov and Mikhailova in 1983 however this view is not accepted by most authors.

Nostoceratidae Extinct family of ammonites

Nostoceratidae is a diverse family of heteromorph ammonites found throughout the oceans of the world during the Late Cretaceous. The nostoceratids are famous for the bizarre coiling of their shells. Many genera, such as Yezoceras, Ainoceras, Anaklinoceras, and some species of Bostrychoceras and Eubostrychoceras, display, as young shells at least, a helical coiling very similar to the shells of the related family, Turrilitidae. As adults, though, the coils then curve away from the axis of coiling, either as an oxbow-like curve around the juvenile coils as in Ainoceras and Anaklinoceras, or in a simple curved loop beneath the juvenile coils, as in Yezoceras. Other genera form loose coils, sometimes in a spiral, such as those of Madagascarites, Muramotoceras, Hyphantoceras, and the infamously convolute Nipponites.

<i>Nostoceras</i> Genus of molluscs (fossil)

Nostoceras is an extinct genus of ammonites. The etymology of the name Nostoceras comes from "nostos" meaning return and "ceros" meaning horn, named as such by Alpheus Hyatt because it bends back on itself.

Ancyloceratoidea, formerly Ancylocerataceae, is a superfamily of typically uncoiled and loosely coiled heteromorph ammonoids established by Alpheus Hyatt in 1900, that may contain as many as 11 families, depending on the classification accepted.

<i>Turrilites</i> Genus of molluscs (fossil)

Turrilites is a genus of helically coiled ammonoid cephalopods from the lower part of the Upper Cretaceous ; generally included in the Ancyloceratina. Previously it was included in the ammonoid suborder, Lytoceratina.

<i>Didymoceras nebrascense</i> Extinct species of ammonite

Didymoceras nebrascense was an extinct species of heteromorph ammonite from the upper Campanian age. It was sexually dimorphic, with two adult sizes averaging at 270 mm (11 in) and 180 mm (7.1 in) high for females and males respectively. It exhibited three distinct growth stages. The first growth stage was composed of one or two straight sharply bending sections and a gently curved third. The second growth stage is composed of around three and a half loosely coiling whorls. The last (adult) growth stage is composed of a U-shaped bend facing upwards.

Turrilitina is a proposed suborder of Cretaceous heteromorph ammonoid cephalopods derived from the Lytoceratidae in the Hauterivian stage of the Lower Cretaceous that would include the superfamilies Turrilitoidea, Ptychoceratoidea, and Scaphatoidea.

Ancyloceratidae Extinct family of ammonites

Ancyloceratidae is a family of heteromorphic ammonites that lived during the Early Cretaceous. Their shells begin as a loose spiral with whorls not touching which then turns into a straight shaft that ends in a J-shape hook or bend at end. Coarse ribbing and spines are common.

<i>Macroscaphites</i>

Macroscaphites is an extinct cephalopod genus included in the Ammonoidea that lived during the Barremian and Aptian stages of the Early Cretaceous. Its fossils have been found throughout most of Europe and North Africa.

Macroscaphitidae Extinct family of ptychoceratoid cephalopods

Macroscaphitidae is an extinct family of ptychoceratoid cephalopods from the subclass Ammonoidea that lived from the Lower Barremian to the Lower Cenomanian stages of the Cretaceous. Fossils of Macroscaphitidae were found all around the world although the abundance of found fossils is rather limited. Known fossils from collections were found largely in Europe, South America and Africa. It is known for some species of which complete specimens were found that these animals developed a hetermorphic shell, i.e. the coiling of the shell was not regular, such that the first whirls formed a planispirally coiled evolute section as seen in homomorphic ammonites, but had an additional straight middle part and a presumably upwards facing aperture. Due to their odd morphology the taxonomic classification of Macroscaphitidae changed often over time since their discovery and may not be finally settled even now.

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