The hellbender, also known as the hellbender salamander, is a species of aquatic giant salamander endemic to the eastern and central United States. It is the largest salamander in North America. A member of the family Cryptobranchidae, the hellbender is the only extant member of the genus Cryptobranchus. Other closely related salamanders in the same family are in the genus Andrias, which contains the Japanese and Chinese giant salamanders. The hellbender is much larger than any other salamander in its geographic range, and employs an unusual adaption for respiration through cutaneous gas exchange via capillaries found in its lateral skin folds. It fills a particular niche — both as a predator and prey — in its ecosystem, which either it or its ancestors have occupied for around 65 million years. The species is listed as Vulnerable on the IUCN Red List of Threatened Species due to the impacts of disease and widespread habitat loss and degradation throughout much of its range.
Amphiuma is a genus of aquatic salamanders from the United States, the only extant genus within the family Amphiumidae. They are colloquially known as amphiumas. They are also known to fishermen as "conger eels" or "Congo snakes", which are zoologically incorrect designations or misnomers, since amphiumas are actually salamanders, and not fish, nor reptiles and are not from Congo. Amphiuma exhibits one of the largest complements of DNA in the living world, around 25 times more than a human.
The Cryptobranchidae are a family of large salamanders that are fully aquatic. The family includes some of the largest living amphibians. They are native to China, Japan, and the eastern United States. Giant salamanders constitute one of two living families—the other being the Asiatic salamanders belonging to the family Hynobiidae—within the Cryptobranchoidea, one of two main divisions of living salamanders.
The Molidae comprise the family of the molas or ocean sunfishes, unusual fish whose bodies come to an end just behind the dorsal and anal fins, giving them a "half-fish" appearance. They are also the largest of the ray-finned bony fish, with the southern sunfish, Mola alexandrini, recorded at 4.6 m (15 ft) in length and 2,744 kg (6,049 lb) in weight.
Peltosaurus is an extinct genus of anguid lizard from North America that lived from the Eocene to the Miocene. Peltosaurus belongs to the anguid subfamily Glyptosaurinae. The type species Peltosaurus granulosus was named in 1873 by American paleontologist Edward Drinker Cope. Many additional species have been named, but most have been reassigned to different genera. For example, Peltosaurus piger, named in 1928, was reclassified as Odaxosaurus piger, and P. jepseni, named in 1942 from the Paleocene of Wyoming, but was later reclassified as Proxestops jepseni. In 1955 a new species, Peltosaurus macrodon, was named from the Eocene of California. Lizard bones from the Late Miocene of Nebraska were attributed to a new species of Peltosaurus called P. minimus in 1976, extending the fossil range of Peltosaurus and Glyptosaurinae into the Neogene. However, these bones were later referred to a genus of skinks called Eumeces, meaning that the fossil range of Peltosaurus and Glyptosaurinae does not go beyond the Paleogene.
The Chinese giant salamander is one of the largest salamanders and one of the largest amphibians in the world. It is fully aquatic, and is endemic to rocky mountain streams and lakes in the Yangtze river basin of central China. It has also been introduced to Kyoto Prefecture in Japan, and possibly to Taiwan. It is considered critically endangered in the wild due to habitat loss, pollution, and overcollection, as it is considered a delicacy and used in traditional Chinese medicine. On farms in central China, it is extensively farmed and sometimes bred, although many of the salamanders on the farms are caught in the wild. It has been listed as one of the top-10 "focal species" in 2008 by the Evolutionarily Distinct and Globally Endangered project.
Rhineura is a genus of worm lizard endemic to North America. The genus has only one extant species but more are known from fossil record. They are also known as the North American worm lizards.
Andrias scheuchzeri is an extinct species of giant salamander belonging to the genus Andrias, which also contains the closely related living Asian giant salamanders. It is known from Oligocene to Pliocene aged deposits primarily from Central Europe, but possibly as far east as Western Siberia and eastern Kazakhstan.
Mourasuchus is an extinct genus of giant, aberrant caiman from the Miocene of South America. Its skull has been described as duck-like, being broad, flat, and very elongate, superficially resembling Stomatosuchus from the Late Cretaceous.
Andrias is a genus of giant salamanders. It includes the largest salamanders in the world, with A. japonicus reaching a length of 1.44 metres, and A. sligoi reaching 1.80 metres. While extant species are only known from East Asia, several extinct species in the genus are known from late Oligocene and Neogene aged fossils collected in Europe and North America, indicating that the genus formerly had a much wider range.
Hesperotestudo is an extinct genus of tortoise native to North and Central America from the Early Miocene to the Late Pleistocene. Species of Hesperotestudo varied widely in size, with a large undescribed specimen from the Late Pleistocene of El Salvador reaching 150 cm (4.9 ft) in carapace length, larger than that of extant giant tortoises. Historically considered a subgenus of Geochelone, it is now considered to be distantly related to that genus. Its relationships with other tortoises are uncertain. The exposed areas of the bodies of Hesperotestudo species were extensively covered with large dermal ossicles, which in life were covered in keratin. It has been suggested that species of Hesperotestudo were relatively tolerant of cold weather. Hesperotestudo became extinct at the end of the Pleistocene roughly co-incident with the arrival of the first humans in North America. There is apparently a site in Florida where one individual may have been killed that some suggested were evidence of butchering, although others suggested that the turtle was neither cooked nor does a ledge that was found near it date at the same time as it.
Lisserpeton is an extinct genus of prehistoric salamander of the Hell Creek Formation. Its closest living relatives are the mole salamanders.
Nordenosaurus is an extinct genus of crocodilian. When first named in 1973 the genus was thought to be a squamate and was assigned to the family Xenosauridae. A single frontal bone was found from the Norden Bridge locality of the lower Valentine Formation in Brown County, Nebraska, thought to date back to the late Miocene. The size of the bone was initially taken as evidence that it was a giant xenosaurid. The specific name of the type species, N. magnus, alludes to its extremely large size in comparison to other xenosaurids known at the time. In 1982, paleontologist Jacques Gauthier reinterpreted Nordenosaurus as a small crocodilian rather than a giant xenosaurid. The frontal bone is hour-glass shaped and covered in deep pits, unlike those of any lizard but similar to the frontals of most crocodilians.
The Ash Hollow Formation of the Ogallala Group is a geological formation found in Nebraska and South Dakota. It preserves fossils dating back to the Neogene period. It was named after Ash Hollow, Nebraska and can be seen in Ash Hollow State Historical Park. Ashfall Fossil Beds State Historical Park is within this formation.
The Valentine Formation is a geologic unit formation or member within the Ogallala unit in northcentral Nebraska near the South Dakota border. It preserves fossils dating to the Miocene epoch of the Neogene period and is particularly noted for Canid fossils. This unit consists of loosely-consolidated sandstone that crumbles easily. These sands carry the water of the Ogallala Aquifer and is the source of much of the water in the Niobrara River. A particular feature of the Valentine is lenticular beds of green-gray opaline sandstone that can be identified in other states, including South Dakota, Nebraska, Kansas, and Colorado. Although three mammalian fauna stages can be mapped throughout the range of the Ogallala, no beddings of the Ogallala are mappable and all attempts of formally applying the Valentine name to any mappable lithology beyond the type location have been abandoned. Even so, opaline sandstone has been used to refer to this green-gray opalized conglomerate sandstone that is widely found in the lower Ogallala Formation.
The Marsland Formation is a geologic formation in Nebraska. It preserves fossils dating back to the Neogene period.
Necturus krausei is an extinct species of mudpuppy salamanders from the Paleocene of Saskatchewan in Canada. It is known from a set of vertebrae found in the Ravenscrag Formation.
The Ozark hellbender is a subspecies of the hellbender. The subspecies is strictly native to the mountain streams of the Ozark Plateau in southern Missouri and northern Arkansas. Its nicknames include lasagna lizard and snot otter. This large salamander grows to a total length of 29–57 cm (11–22 in) over a lifespan of 30 years. The Ozark hellbender is a nocturnal predator that hides under large flat rocks and primarily consumes crayfish and small fish. As of 2011, the U.S. Fish and Wildlife Service (USFWS) has listed the subspecies as endangered under the Endangered Species Act of 1973. The population decline of the subspecies is caused by habitat destruction and modification, overutilization, disease and predation, and low reproductive rates. Conservation programs have been put in place to help protect the subspecies.
Maliamia is an extinct genus of amiid ray-finned fish from the Early Eocene, known from fragmentary remains found in the Tamaguélelt Formation of Mali. It was described in 1989, based on fossils recovered by three separate expeditions in 1975, 1979–80, and 1981. The type species is Maliamia gigas, named in reference to its large size.
Sacacosuchus is an extinct monospecific genus of marine gavialid that lived along the coast of the south-east Pacific from approximately 19 to 6.3 million years ago. Its fossils have been found in the Chilcatay and Pisco Formations of Peru, where it coexisted with the much larger Piscogavialis. Based on its skull, Sacacosuchus was most likely a generalist feeder with an estimated total body length of 4.32 m (14.2 ft). Its extinction is thought to have been caused by a combination of factors including falling sea levels and global cooling.
