Belinae | |
---|---|
Rhicnobelus metallicus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Infraorder: | Cucujiformia |
Family: | Belidae |
Subfamily: | Belinae Schönherr, 1826 |
Tribes | |
Belinae are a Gondwanan subfamily of beetles which belong to the belids, primitive weevils of the family Belidae. Like in other belids, their antennae are straight, not elbowed as in the true weevils (Curculionidae). The Belinae make up the bulk of the diversity of living belid genera. They are found in the Australia-New Guinea-New Zealand and South America.
Of the three tribes placed here, the Pachyurini appear to hold the most primitive genera, while the Agnesiotidini might be the most advanced one. The overall delimitation of tribes is not entirely robust though, and particularly the placement of the diverse Belini in respect to the Agnesiotidini is somewhat uncertain. The belids as a whole are of Jurassic origin, and the Belinae must thus date from the Late Jurassic or at most Early Cretaceous, roughly some 150 million years ago. [1]
Belinae typically have an elongated and cylindrical shape. They can be distinguished from the Oxycoryninae (including the former Aglycyderinae) by a few characters: The scutellum of the Belinae is tipped upwards, with the base hidden by a flange at the elytra bases and the distal end pointing out between the wing bases. The sternite of the metathorax is characteristically swollen. Microscopically, it can be seen that their spermatheca are sickle-shaped, well developed and darkly pigmented. The Agnesiotidini have a groove at the hind margin of the eye, which is missing in the other two tribes. [2]
In the larvae, the posterior margin of the pronotum is enlarged, with a more (Agnesiotidini and Belini) or less (Pachyurini) strongly thickened hind margin. In the Pachyurini, the larvae has an open and divided occipital foramen with ridges on the dorsal emargination and the labrum and epipharynx have paired and divergent paramesal sclerotizations. These are also present but curved in the Agnesiotidini; the labrum-epipharynx of the Belini is unsclerotized. The maxillary malae are densely covered with setae in the Agnesiotidini, it has only a few setae in the others. [2]
Their larvae feed on wood, mostly of Araucariaceae. Certain Belini larvae also feed on the wood of Fabaceae, while some Agnesiotidini take the wood of Myrtaceae. Some Pachyurini larvae eat the developing fruits of Nothofagaceae. Like all Belini, they prefer diseased or dying plants or deadwood to healthy plants. [2]
The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families with 6,800 genera and 83,000 species described worldwide. They are the sister group to the family Brentidae.
Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small–less than 6 mm in length–and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, with most of them in the family Curculionidae. It also includes bark beetles, which while morphologically dissimilar to other weevils in lacking the distinctive snout, is a subfamily of Curculionidae. Some other beetles, although not closely related, bear the name "weevil", such as the biscuit weevil, which belongs to the family Ptinidae.
Darkling beetle is the common name for members of the beetle family Tenebrionidae. The number of species in the Tenebrionidae is estimated at more than 20,000 and the family is cosmopolitan in distribution.
Amphizoa is a genus of aquatic beetles in the suborder Adephaga, placed in its own monogeneric family, Amphizoidae. There are five known species of Amphizoa, three in western North America and two in the eastern Palearctic. They are sometimes referred to by the common name troutstream beetles.
The beetle subfamily Curculioninae is part of the weevil family Curculionidae. It contains over 23,500 described species in 2,200 genera, and is therefore the largest weevil subfamily. Given that the beetle order (Coleoptera) contains about one-quarter of all known organisms, the Curculioninae represent one of the – if not the – most successful radiations of terrestrial Metazoa.
Belidae is a family of weevils, called belids or primitive weevils because they have straight antennae, unlike the "true weevils" or Curculionidae which have geniculate (elbowed) antennae. They are sometimes known as "cycad weevils", but this properly refers to a few species from the genera Parallocorynus and Rhopalotria.
Hybosoridae, sometimes known as the scavenger scarab beetles, is a family of scarabaeiform beetles. The 690 species in 97 genera occur widely in the tropics, but little is known of their biology.
Brentidae, sometimes known as the primitive weevils, is a cosmopolitan family of primarily xylophagous beetles also known as straight-snouted weevils. The concept of this family has been expanded with the inclusion of three groups formerly placed in the Curculionidae; the subfamilies Apioninae, Cyladinae, and Nanophyinae, as well as the Ithycerinae, previously considered a separate family. They are most diverse in the tropics, but occur throughout the temperate regions of the world. They are among the families of weevils that have non-elbowed antennae, and tend to be elongate and flattened, though there are numerous exceptions.
The Chrysomelinae are a subfamily of leaf beetles (Chrysomelidae), commonly known as broad-bodied leaf beetles or broad-shouldered leaf beetles. It includes some 3,000 species around the world.
Anthribidae is a family of beetles also known as fungus weevils. The antennae are not elbowed, may occasionally be longer than the body and thread-like, and can be the longest of any members of Curculionoidea. As in the Nemonychidae, the labrum appears as a separate segment to the clypeus, and the maxillary palps are long and projecting.
The Attelabidae is a widespread family of weevils. They are among the primitive weevils, because of their straight antennae, which are inserted near the base of the rostrum. The prothorax is much narrower than the base of the elytra on the abdomen. Attelabidae and the related family Rhynchitidae are known commonly as the leaf-rolling weevils. Rhynchitidae may be treated as subfamily Rhynchitinae of the Attelabidae.
Nemonychidae is a small family of weevils, placed within the primitive weevil group because they have straight rather than geniculate (elbowed) antennae. They are often called pine flower weevils. As in the Anthribidae, the labrum appears as a separate segment to the clypeus, and the maxillary palps are long and projecting. Nemonychidae have all ventrites free, while Anthribidae have ventrites 1-4 connate or partially fused. Nemonychidae lack lateral carinae on the pronotum, while these are usually present, though may be short, in Anthribidae.
Jubaeopsis caffra, the Pondoland palm, is a flowering plant species in the palm family (Arecaceae). It belongs to the monotypic genus Jubaeopsis.
The dustywings, Coniopterygidae, are a family of Pterygota of the net-winged insect order (Neuroptera). About 460 living species are known. These tiny insects can usually be determined to genus with a hand lens according to their wing venation, but to distinguish species, examination of the genitals by microscope is usually necessary.
Eobelinae are an extinct weevil subfamily that throve in the late Mesozoic. They belong to the family Nemonychidae. The Eobelidae were widespread, occurring at least in Central Asia, Spain and Brazil during the Late Jurassic and Early Cretaceous.
Oxycoryninae are subfamily of primitive weevils of the family Belidae, but sometimes treated as a distinct family Oxycorynidae. Like in other belids, their antennae are straight, not elbowed as in the true weevils (Curculionidae), and their larvae feed on the wood of diseased or dying plants or on deadwood or fruits; they tend to avoid healthy plants.
Aglycyderini is a tribe of belids, primitive weevils of the family Belidae. Like in other belids, their antennae are straight, not elbowed as in the true weevils (Curculionidae). They occur only on the Pacific Islands and in the Macaronesian region.
Metrioxenini (metrioxenines) are a tribe of belids, primitive weevils of the family Belidae, containing about 30 species. They are only found in two widely separated areas, Southeast Asia extending to Indonesia, and South Africa. In the Paleogene, they were found at least in North America and Europe also, occurring perhaps across the entire Northern Hemisphere.
Oxycorynini are a tribe of American belids, primitive weevils of the family Belidae. Like in other belids, their antennae are straight, not elbowed as in the true weevils (Curculionidae). In contrast to other Belidae, Oxycorynini larvae feed on flowers or several types of fruit.
Naupactini is a tribe of broad-nosed weevils. Primarily from the Neotropical realm, reaches highest genus and species diversity in South America. Their size varies from 3.5 to 35 mm long, and its colour patterns are diverse. As well many has colourful iridescent scales, others show opaque scales or setae, and some are subglabrous. In habitats with sparse vegetation or trees absence, the occurrence of flightlessness and parthenogenesis is frequent.