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Fossil of Beloceras sagittarium | |
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Family: | Beloceratidae Hyatt 1884 |
Beloceratidae is a family of ammonites included in the order Agoniatitida.
These fast-moving nektonic carnivores lived in the Late Devonian period, from 379.5 to 376.1 Ma.
Fossils of species within this genus have been found in the Devonian sediments of Algeria, France, Germany, Morocco, Spain and United States. [1]
The Devonian is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.2 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.9 Ma. It is named after Devon, South West England, where rocks from this period were first studied.
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Sarcopterygii — sometimes considered synonymous with Crossopterygii — is a clade of bony fish commonly referred to as lobe-finned fish. They are characterised by prominent muscular limb buds (lobes) within their fins, which are supported by articulated appendicular skeletons. This is in contrast to the other clade of bony fish, the Actinopterygii, which have only skin-covered bony spines supporting the fins.
Placoderms are members of the class Placodermi, a group of prehistoric fish known from Paleozoic fossils which lived from the Silurian to the end of the Devonian period. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates, and the rest of the body was scaled or naked depending on the species. Placoderms were among the first jawed fish; their jaws likely evolved from the first pair of gill arches.
The Late Devonian extinction consisted of several extinction events in the Late Devonian Epoch, which collectively represent one of the five largest mass extinction events in the history of life on Earth. The term primarily refers to a major extinction, the Kellwasser event, also known as the Frasnian-Famennian extinction, which occurred around 372 million years ago, at the boundary between the Frasnian stage and the Famennian stage, the last stage in the Devonian Period. Overall, 19% of all families and 50% of all genera became extinct. A second mass extinction called the Hangenberg event, also known as the end-Devonian extinction, occurred 359 million years ago, bringing an end to the Famennian and Devonian, as the world transitioned into the Carboniferous Period.
The Givetian is one of two faunal stages in the Middle Devonian Period. It lasted from 387.7 million years ago to 382.7 million years ago. It was preceded by the Eifelian Stage and followed by the Frasnian Stage. It is named after the town of Givet in France. The oldest forests occurred during the late Givetian. The lower GSSP is located at Jebel Mech Irdane, Tafilalt, Morocco.
The Frasnian is one of two faunal stages in the Late Devonian Period. It lasted from 382.7 million years ago to 372.2 million years ago. It was preceded by the Givetian Stage and followed by the Famennian Stage.
The Famennian is the later of two faunal stages in the Late Devonian epoch. The most recent estimate for its duration is that it lasted from around 371.1 to 359.3 million years ago. An earlier 2012 estimate, still used by the International Commission on Stratigraphy, is that it lasted from 372.2 million years ago to 358.9 million years ago. It was preceded by the Frasnian stage and followed by the Tournaisian stage.
Arthrodira is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of placoderms.
The Emsian is one of three faunal stages in the Early Devonian Epoch. It lasted from 407.6 ± 2.6 million years ago to 393.3 ± 1.2 million years ago. It was preceded by the Pragian Stage and followed by the Eifelian Stage. It is named after the Ems river in Germany. The GSSP is located in the Zinzil'ban Gorge in the Kitab State Geological Reserve of Uzbekistan, 35 centimetres (14 in) above the contact with the Madmon Formation.
The Lochkovian is one of three faunal stages in the Early Devonian Epoch. It lasted from 419.2 ± 3.2 million years ago to 410.8 ± 2.8 million years ago. It marked the beginning of the Devonian Period, and was followed by the Pragian Stage. It is named after the village of Lochkov in the Czech Republic, now part of the city of Prague. The GSSP is located within the Lochkow Formation at the Klonk Section in Prague.
The Early Devonian is the first of three epochs comprising the Devonian period, corresponding to the Lower Devonian series. It lasted from 419.2 ± 3.2 to 393.3 ± 1.2 and began with the Lochkovian Stage 419.2 ± 3.2 to 410.8 ± 1.2, which was followed by the Pragian from 410.8 ± 3.2 to 407.6 ± 1.2 and then by the Emsian, which lasted until the Middle Devonian began, 393.3± 1.2 million years ago. During this time, the first ammonoids appeared, descending from bactritoid nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era.
The Orcadian Basin is a sedimentary basin of Devonian age that formed mainly as a result of extensional tectonics in northeastern Scotland after the end of the Caledonian orogeny. During part of its history, the basin was filled by a lake now known as Lake Orcadie. In that lacustrine environment, a sequence of finely bedded sedimentary rocks was deposited, containing well-preserved fish fossils, with alternating layers of mudstone and coarse siltstone to very fine sandstone. These flagstones split easily along the bedding and have been used as building material for thousands of years. The deposits of the Orcadian Basin form part of the Old Red Sandstone (ORS). The lithostratigraphic terms lower, middle and upper ORS, however, do not necessarily match exactly with sediments of lower, middle and upper Devonian age, as the base of the ORS is now known to be in the Silurian and the top in the Carboniferous.
Elpistostegalia is a clade containing Panderichthys and all more derived tetrapodomorph taxa. The earliest elpistostegalians, combining fishlike and tetrapod-like characters, such as Tiktaalik, are sometimes called fishapods. Although historically Elpistostegalia was considered an order of prehistoric lobe-finned fishes, it was cladistically redefined to include tetrapods.
The geology of the Orkney islands in northern Scotland is dominated by the Devonian Old Red Sandstone (ORS). In the southwestern part of Mainland, this sequence can be seen to rest unconformably on a Moinian type metamorphic basement.
The Rhynchonellata is a class of Lower Cambrian to Recent articulate brachiopods that combines orders from within the Rhynchonelliformea with well developed pedicle attachment. Shell forms vary from those with wide hinge lines to beaked forms with virtually no hinge line and from generally smooth to strongly plicate. Most all are biconvex. Lophophores vary and include both looped and spiraled forms. Although morphologically distinct, included orders follow a consistent phylogenetic sequence.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
Antarctilamna is an extinct genus of Devonian cartilaginous fish originally exemplified by Antarctilamna prisca from South Eastern Australia and Antarctica. The latest occurring described species is Antarctilamna ultima from the Waterloo Farm lagerstätte in South Africa. Antarctilamna has robust ctenacanthid-like spines which lack a deep insertion area, and are borne in front of the first dorsal fin; in addition to distinctive diplodont teeth with small intermediate cusps. Antarctilamna-like spines, known from the Bunga Beds locality in Australia have been ascribed to A. prisca.
Tutusius is a genus of extinct tetrapod from the Devonian of South Africa, containing a single species, Tutusius umlambo. It was described from the +/- 360 myo Gondwana locality of Waterloo Farm lagerstätte on the south-eastern coast of South Africa, which at the time was located within the Antarctic Circle. Together with the find of Umzantsia amazana from the same locality, this provides the first evidence that Devonian tetrapods were not restricted to the tropics as was formerly believed, and suggests that they may have been global in distribution. Waterloo Farm fossils have been metamorphosed and intensely flattened, with the bone tissue replaced by secondary metamorphic mica that is partially altered to kaolinite and chlorite during uplift. They also provide the first evidence of Devonian tetrapods from the continent of Africa, and only the second and third such taxa from Gondwana.
Umzantsia is an extinct genus of limbed stegocephalians from the Devonian of South Africa. It contains a single species, Umzantsia amazana. The genus is based on ~360 million year old skull and shoulder bones from the Waterloo Farm lagerstätte, a locality on the south-eastern coast of South Africa. In the Devonian, this area of Gondwana was located within the Antarctic Circle. Together with the find of Tutusius umlambo from the same locality, this provides the first evidence that Devonian limbed vertebrates were not restricted to the tropics as was formerly believed, and suggests that they may have been global in distribution. Waterloo Farm fossils have been metamorphosed and intensely flattened, with the bone tissue replaced by secondary metamorphic mica that is partially altered to kaolinite and chlorite during uplift. They also provide the first evidence of Devonian limbed vertebrates from the continent of Africa, and only the second and third such taxa from Gondwana.