Cassianellidae

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Cassianellidae
Temporal range: Landinian - Norian 237.0–203.6  Ma
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Mollusca
Class: Bivalvia
Order: Pteriida
Superfamily: Pterioidea
Genus: Cassianellidae
Ichikawa, 1958
Genera

See text

Cassianellidae is a small extinct family of prehistoric bivalves which lived from the Middle Triassic, Ladinian stage through the Late Triassic Norian stage. [1] It has been suggested that the family may have evolved from the related family Bakevelliidae. [2] Bakevelliidae species are found in North America, South America, Europe, the Middle East, and Asia. Living a stationary life attached to substrate in marine and brackish environments, they formed shells of an aragonite composition with a low amount of magnesium calcite. The family Cassianellidae was named by K. Ichikawa in 1958. [1]

Contents

Description

The cassianellids are noted for being restricted to the Triassic only, going extinct during the end Triassic extinction event. They are thought to have a close relationship to the longer lived family Bakevelliidae and it is possible the cassianellids may have even evolved from one of the early Triassic bakevelliid genera. [2] As is found in the Bakevelliidae genus Kobayashites and the Isognomonidae genus Lithiotis , cassianellids are distinguished by the presence of a larval septum which is retained in the adult shell morphology. [2] Also notable for the entire family is the presence of a primitive alivincular type ligament, termed an alivincular-areate ligament. This form of ligament was widely found in many of the Triassic bivalve families but evolution of more refined ligament systems resulted in the shift away from the alivincular-areate system. Only a few modern families, such as Gryphaeidae, still retain the alivincular-areate ligament system. [3] Several of the genera, such as Cassianella , have a widespread distribution. [1] In contrast, Lilangina is known only from Asia, and Septihoernesia from the middle Triassic of northern Italy. [1]

Taxonomy

CassianellidaeIchikawa, 1958

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The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 541 million years to the present, and began with the Cambrian Period when animals first developed hard shells preserved in the fossil record. The time before the Phanerozoic, called the Precambrian, is now divided into the Hadean, Archaean and Proterozoic eons.

Triassic First period of the Mesozoic Era 252-201 million years ago

The Triassic is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.9 million years ago (Mya), to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic.

Bivalvia Class of molluscs

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Nautiloid subclass of molluscs

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Placodontia Extinct order of marine reptiles

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Pteriida order of molluscs

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Trigoniidae family of molluscs

Trigoniidae is a taxonomic family of saltwater clams, marine bivalve mollusks in the superfamily Trigonioidea. There is only one living genus, Neotrigonia, but in the geological past this family was well represented, widespread and common. The shells of species in this family are morphologically unusual, with very elaborate hinge teeth, and the exterior of the shell is highly ornamented.

Gryphaeidae family of molluscs

The Gryphaeidae, common name the foam oysters or honeycomb oysters, are a family of marine bivalve mollusks, and are a kind of true oyster. This family of bivalves is very well represented in the fossil record, however the number of living species is very few.

Mesozoic marine revolution

The Mesozoic Marine Revolution (MMR) is the rapid adaption to shell-crushing (durophagous) and boring predation in benthic organisms throughout the Mesozoic era. The term was first coined by Geerat J. Vermeij, who based his work on that of Steven M. Stanley. While initially restricted to the Late Cretaceous, more recent studies have suggested that the beginning of this ecological arms race extends back into the Norian It is the important transition between the Palaeozoic evolutionary fauna and the Cenozoic evolutionary fauna that occurred throughout the Mesozoic.

Hybodontiformes order of fishes (fossil)

Hybodontiformes, also called hybodonts, are an extinct subset of Elasmobranchii which existed from the Devonian to the Miocene. They form the group of sharks closest to neoselachians, the clade of modern sharks and rays. Hybodonts were named and are distinguished based on their conical tooth shape. They comprised the main group of Jurassic sharks in Europe and North America. They survived into the Late Cretaceous before going extinct, possibly due to competition from other sharks, though forms like Miosynechodus endured as recently as the Miocene. Lonchidion was one of the last hybodonts — its distinctive serrated fine spines occur in freshwater deposits from Wyoming alongside the fossils of the last dinosaurs, including Tyrannosaurus rex and Triceratops. Hybodontiformes are identified in the fossil record predominantly based on distinct teeth and fin spines. They were known to live in both fresh and salt water environments.

Prolecanitida is an order of extinct ammonoid cephalopods with discoidal to thinly lenticular shells with goniatitic or ceratitic sutures and which retained the simple retrochoanitic siphuncle with backward extending septal necks. As typical for ammonoids the siphuncle is along the ventral margin. Prolecanitids form a relatively small and stable order within the Ammonoidea with 43 named genera and about 1250 species, but with a long-ranging lineage of about 108 m.y. stretching from the Lower Carboniferous to the Triassic. Although not as diverse as their goniatitid contemporaries, the Prolecanatida provided the stock from which all later Mesozoic ammonoids were derived.

Pojetaia is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early to Middle Cambrian fossils found in North America, Greenland, Europe, North Africa, Asia, and Australia. The genus currently contains two accepted species, Pojetaia runnegari, the type species, and Pojetaia sarhroensis, though up to seven species have been proposed. The genera Buluniella, Jellia, and Oryzoconcha are all considered synonyms of Pojetaia.

Bakevelliidae family of molluscs (fossil)

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Fordilloidea superfamily of molluscs (fossil)

Fordilloidea is an extinct superfamily of early bivalves containing two described families, Fordillidae and Camyidae and the only superfamily in the order Fordillida. The superfamily is known from fossils of early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia. Fordillidae currently contains two genera, Fordilla and Pojetaia each with up to three described species while Camyidae only contains a single genus Camya with one described species, Camya asy. Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish Small shelly fauna.

References

  1. 1 2 3 4 The Paleobiology Database Cassianellidae entry accessed 2 January 2012
  2. 1 2 3 Malchus, N. (2004). "Early ontogeny of Jurassic bakevelliids and their bearing on bivalve evolution" (PDF). Acta Palaeontologica Polonica. 49 (1): 85–110.
  3. Hautmann, M. (2004). "Early Mesozoic evolution of alivincular bivalve ligaments and its implications for the timing of the 'Mesozoic marine revolution'" (PDF). Lethaia. 37: 165–172. doi:10.1080/00241160410005835. Archived from the original (PDF) on 2012-04-07.