| Chamaesomatidae | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Subphylum: | Myriapoda |
| Class: | Diplopoda |
| Order: | Chordeumatida |
| Superfamily: | Brannerioidea |
| Family: | Chamaesomatidae |
Chamaesomatidae is a family of millipedes belonging to the order Chordeumatida. [1] This family contains 28 species distributed among 11 genera. These millipedes are found in Europe and North Africa. [2]
These millipedes range from 3.3 mm to 12 mm in length. Many species dwell in caves. The paranota are small and humplike. Adult millipedes in this family have 26, 28, or 30 segments (counting the collum as the first segment and the telson as the last). [2] This family includes the species Chamaesoma broelemanni , notable as one of only a few chordeumatidan species with only 26 segments in adults, four fewer segments than typically found in adults in this order. [3] [4] Most genera in this family lack pigment. [5]
Adult males in this family feature anterior gonopods derived from the eighth leg pair and posterior gonopods derived from the ninth leg pair. The anterior gonopods each include a distal element, the telopodite, which is usually well developed, and a basal element, the angiocoxite, which is derived from the base (coxa) of the eighth leg pair, as well as another structure, the colpocoxite, which is thought to be permanently extruded and sclerotized walls of the coxal gland. [2] [5] [6] The anterior gonopods sometimes feature flagella. The posterior gonopods each feature a telepodite with either one or two segments. The eleventh pair of legs in adult males usually feature coxal hooks. [2] [5]
This family includes 11 genera: [1]
Chordeumatida is a large order of millipedes containing more than 1,400 species. Also known as sausage millipedes, they are found nearly worldwide. Chordeumatida is the largest order in the superorder Nematophora, a group also known as spinning millipedes because their telsons feature spinnerets used to build nests of silk. These millipedes produce this silk to create chambers in which to molt or to lay their eggs.
Haplodesmidae is a family of millipedes in the order Polydesmida. This family includes about 70 species. Species occur in East Asia, Southeast Asia, and Oceania, although some species have been introduced to the New world tropics.
Glomeridesmidae is a millipede family of the order Glomeridesmida. This family includes two genera: The genus Glomeridesmus includes most species in this family; the genus Glomeridesmoides includes one species.
Anthroleucosomatidae is a family of millipedes in the order Chordeumatida. This family includes more than 100 species distributed among 40 genera. This family has a mostly Holarctic distribution, with the greatest diversity in the area around the Mediterranean sea.
Craspedosomatidae is a family of millipedes in the order Chordeumatida. Most adult millipedes in this family have 30 segments, but some have only 28. There are at least 30 genera and 210 described species in Craspedosomatidae.
Metopidiotrichidae is a family of millipedes in the order Chordeumatida. This family includes more than 70 species. These millipedes are found in Indochina, Australia, and on Pacific islands from New Zealand to Japan.
Chordeumatidae is a family of millipedes belonging to the order Chordeumatida. These millipedes range from 7 mm to 18 mm in length and are found in Europe. Adult millipedes in this family have either 28 or 30 segments. This family features distinctive sex-linked modifications to the legs in adults: In the adult female, a legless sternite replaces the third pair of legs, and in the adult male, five pairs of legs are modified in the gonopod complex. These modifications are more extensive than those found in other adult males in this order, which often have only two leg pairs modified into gonopods. With the more extensive modifications to the legs in this family, species with the usual 30 segments feature adult females with only 49 leg pairs and adult males with only 45 pairs of walking legs, and species with only 28 segments feature adult females with only 45 leg pairs and adult males with only 41 pairs of walking legs.
Peterjohnsiidae is a small family of millipedes belonging to the order Chordeumatida. The family was first described in 1987 by Jean-Paul Mauriès. These millipedes range from 3 mm to 8 mm in length and are found in Australia. Species in this family exhibit sexual dimorphism in segment number: adult males have 30 segments, but adult females have 32 segments. In adult males in this family, the gonopod complex involves three leg pairs rather than just the two usually modified into gonopods in this order.
Entomobielziidae is a family of millipedes belonging to the order Chordeumatida. This family includes 18 species, including 13 in the genus Tianella. Millipedes in this family are found in Romania, central Asia, and the Himalayas.
Opisthocheiridae is a family of millipedes belonging to the order Chordeumatida. This family includes 40 species distributed among seven genera. These millipedes are found in western Europe and Morocco.
Glomeridellidae is a family of millipedes belonging to the order Glomerida. The adult males in this family have two pairs of telopods rather than the single pair found in other families in this order.
Opisthocheiron canayerensis is a species of millipede in the family Opisthocheiridae. This species is found in France, far inside deep caves. This millipede is notable as one of only a few species in the order Chordeumatida with only 26 segments in adults, four fewer segments than typically found in adults this order.
Chamaesoma is a monotypic genus of millipede in the family Chamaesomatidae, and Chamaesoma broelemanni is the only species in this genus. This millipede is notable as one of only a few species in the order Chordeumatida with only 26 segments in adults, four fewer segments than typically found in adults this order. This species lives on the surface of the soil and is distributed across most of France and Luxembourg. The genus Chamaesoma and its only species were first described by the German zoologist Karl W. Verhoeff in 1913 based on type material found by the French zoologists Henri Brölemann and Henri Ribaut under wet leaf litter near St. Béat in the Pyrenees in France.
Haasea is a genus of millipedes in the family Haaseidae. This genus is the largest in this family, with 17 accepted species. Millipedes in this genus have either 28 or 30 segments.
Haasea hungarica is a species of millipede in the family Haaseidae. This species is a eutroglophile, capable of spending its entire life cycle in a cave but also capable of living on the surface outside of caves. Although these millipedes are often found in caves, this species has also been found on the soil surface in forests.
Lipseuma is a genus of millipedes in the family Kashmireumatidae. This genus contains only two species, the type species L. josianae and its close relative L. bernardi. Both species are troglobites found in caves in China.
Metamastigophorophyllon is a genus of millipedes in the family Anthroleucosomatidae. This genus includes millipedes found in Azerbaijan, Iran, Russia, and Georgia. All five species in this genus have 31 segments in adults. This genus is notable for including the only species in the order Chordeumatida with 31 segments rather than the 30 segments usually observed in adults in this order.
Neocambrisoma is a genus of millipedes in the family Metopidiotrichidae. Millipedes in this genus are found in Tasmania and New South Wales in Australia. Like other genera in this family, this genus features 32 segments in adults of both sexes, rather than the 30 segments usually observed in adults in the order Chordeumatida. Accordingly, female adults in this genus have 54 pairs of legs, which is not only the maximum number observed in this order but also the maximum number fixed by species in the class Diplopoda.
Neocambrisoma raveni is a species of millipede in the family Metopidiotrichidae. These millipedes are found in New South Wales in Australia. Like other species in this family, N. raveni features 32 segments in adults of both sexes, rather than the 30 segments usually observed in adults in the order Chordeumatida. Accordingly, adult females of this species have 54 pairs of legs, which is not only the maximum number observed in this order but also the maximum number fixed by species in the class Diplopoda.