Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta, also known as Coniferophyta or Coniferae. The division contains a single extant class, Pinopsida. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas-firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. The division Pinophyta contains seven families, 60 to 65 genera, and more than 600 living species.
A spruce is a tree of the genus Picea, a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. Picea is the sole genus in the subfamily Piceoideae. Spruces are large trees, from about 20 to 60 m tall when mature, and have whorled branches and conical form. They can be distinguished from other members of the pine family by their needles (leaves), which are four-sided and attached singly to small persistent peg-like structures on the branches, and by their cones, which hang downwards after they are pollinated. The needles are shed when 4–10 years old, leaving the branches rough with the retained pegs. In other similar genera, the branches are fairly smooth.
Picea abies, the Norway spruce or European spruce, is a species of spruce native to Northern, Central and Eastern Europe.
The Tortricidae are a family of moths, commonly known as tortrix moths or leafroller moths, in the order Lepidoptera. This large family has over 11,000 species described, and is the sole member of the superfamily Tortricoidea, although the genus Heliocosma is sometimes placed within this superfamily. Many of these are economically important pests. Olethreutidae is a junior synonym. The typical resting posture is with the wings folded back, producing a rather rounded profile.
Picea glauca, the white spruce, is a species of spruce native to the northern temperate and boreal forests in the United state of North and South, east, and west America. Picea glauca is native from central Alaska all through the east, across western and southern/central Canada to the Avalon Peninsula in Newfoundland, and south to Montana, North Dakota, Minnesota, Wisconsin, Michigan, Upstate New York and Vermont, along with the mountainous and immediate coastal portions of New Hampshire and Maine, where temperatures are just barely cool and moist enough to support it. There is also an isolated population in the Black Hills of South Dakota and Wyoming. It is also known as Canadian spruce, skunk spruce, cat spruce, Black Hills spruce, western white spruce, Alberta white spruce, and Porsild spruce.
Choristoneura fumiferana, the eastern spruce budworm, is a species of moth of the family Tortricidae native to the eastern United States and Canada. The caterpillars feed on the needles of spruce and fir trees. Eastern spruce budworm populations can experience significant oscillations, with large outbreaks sometimes resulting in wide scale tree mortality. The first recorded outbreaks of the spruce budworm in the United States occurred in about 1807, and since 1909 there have been waves of budworm outbreaks throughout the eastern United States and Canada. In Canada, the major outbreaks occurred in periods circa 1910–20, c. 1940–50, and c. 1970–80, each of which impacted millions of hectares of forest. Longer-term tree-ring studies suggest that spruce budworm outbreaks have been recurring approximately every three decades since the 16th century, and paleoecological studies suggest the spruce budworm has been breaking out in eastern North America for thousands of years.
Hylobius warreni, Warren's rootcollar weevil, is a common pest of spruce and pine throughout Canada. It causes considerable damage to native species of spruce both in natural stands and in plantations.
Archips oporana, also known as the pine tortrix or spruce tortrix is a moth of the family Tortricidae, found in Asia and Europe. It was first described by Carl Linnaeus in 1758.
Epinotia nanana, the European spruce needleminer, is a moth of the family Tortricidae. It is found from northern and central Europe to Russia and Mongolia.
Cydia illutana is a small moth of the family Tortricidae. It is found from western and central Europe, north to Scandinavia and east to Russia (Siberia).
Dendroctonus rufipennis, the spruce beetle, is a species of bark beetle native to British Columbia, Newfoundland and Labrador, Nova Scotia, Ontario, Quebec, Northern Manitoba, the Yukon, Alaska, Colorado, Wyoming, Montana, and Maine. They are known to destroy forests of spruce trees including Engelmann, White, Sitka, and Colorado blue spruce. Adults average 4 to 7 mm in length.
Coleotechnites piceaella is a moth of the family Gelechiidae. It is found in north-eastern United States and Canada. It is an introduced species in Europe and was first recorded from Great Britain in 1952, then Germany in 1963 and has expanded from there over all of central Europe, towards France, Italy and Hungary.
Synanthedon pini, the pitch mass borer, is a moth of the family Sesiidae. The pitch mass borer occurs on spruce and pine in eastern North America. It does not kill trees, but the pitch-filled larval tunnels in the wood cause defects in the lumber.
Dioryctria reniculelloides, the spruce coneworm, is a moth of the family Pyralidae. The species was first described by Akira Mutuura and Eugene G. Munroe in 1973. It is found from Nova Scotia to Alaska, south in the east to New York, and south in the west to California and New Mexico. It was recorded from China in 2009. Occasionally abundant, often in conjunction with epidemics of the spruce budworm, the spruce coneworm occurs through most or all of the range of spruce in North America, feeding on new foliage and cones of spruce, and often balsam fir. When abundant, it can be a serious pest "particularly on white spruce".
Argyresthia picea is a moth of the family Yponomeutidae. It is found in Canada, including Alberta, Ontario, Quebec and the Yukon.
Archips packardianus, the spring spruce needle moth or spruce needleworm, is a species of moth of the family Tortricidae. Spruce needle worms are commonly found in small numbers on spruce and trees of other coniferous genera throughout most of Canada and the northeastern US. Archips packardiana overwinters as a tiny larva in a mined needle. Needle mining is resumed in the spring, the larvae later moving to feed on new foliage, where they spin considerable webbing. Full-grown larvae have a pale green head, sometimes patterned with brown, and a pale body and pale thoracic legs, and are about 20 mm long. The larva pupates, usually in the webbed needles, and the adult emerges in summer to early fall. The closely related Archips strianus is much less common, but probably has a habitat similar to that of A. packardiana. The larvae have dark thoracic legs and small dark areas around the base of the thoracic hairs. The habitat consists of coniferous and mixed woods.
Megastigmus atedius is a species of minute wasp that feeds on white spruce seed and cones. The damage it causes is largely undetected because the larvae complete their development hidden inside the seeds, which reveal no external indication of this. Although species of Megastigmus are said to be host-specific, the spruce seed chalcid found near Fairbanks, Alaska, was identified by E.H. Holsten and others in 1980 as M. piceae, while A.H. Rose and O.H. Lindquist applied the name Megastigmus piceae, but gave the authority as Rohwer.
Barbara mappana is a species of moth in the family Tortricidae. It has been reported on cones of white spruce, but is also considered to be of minor importance compared to other insect species which consume and potentially damage white spruce.
Gilpinia hercyniae, the European spruce sawfly, was first found in North America near Hull, Quebec, in 1922. By 1930, severe damage was occurring in spruce stands in the Lower St. Lawrence–Gaspé region of Quebec. Population levels exploded, and further damage subsequently occurred in New Brunswick and the northeastern United States. Loss of wood from spruce mortality was estimated at 10 million cubic metres, but by the mid-1980s pest numbers on all species of spruce in Canada from Manitoba to the Atlantic Ocean and in adjacent parts of the United States had become relatively low under the influence of spontaneous virus disease and introduced parasites. In Newfoundland, control was helped by the introduction of the masked shrew, a cocoon-hunting insectivore.
Spruce sawflies are various sawfly species found in North America that feed on spruce. There are multiple species of sawflies known as spruce sawflies, including species in the genera Gilpinia, Pikonema, Pristiphora, and Cephalcia. Each kind of sawfly attacks particular parts of the spruce as larvae during different times of the year.
