Cystopteris is a genus of ferns in the family Cystopteridaceae. These are known generally as bladderferns or fragile ferns. They grow in temperate areas worldwide. This is a very diverse genus and within a species individuals can look quite different, especially in harsh environments where stress stunts their growth. They hybridize easily with each other and identifying an individual can be challenging. In general these are rhizomatous perennials which grow in rocks or soil. Their leaves are multiply pinnate, in that each leaflet is divided into smaller parts. The sori are usually rounded and covered in an inflated bladder-like indusium.
Cystopteris fragilis is a species of fern known by the common names brittle bladder-fern and common fragile fern. It can be found worldwide, generally in shady, moist areas. The leaves are up to 30 or 40 centimeters long and are borne on fleshy petioles with few or no long hairs. Each leaf is divided into many pairs of leaflets, each of which is subdivided into lobed segments. The underside of the leaf has many rounded sori containing the sporangia.
Cystopteris protrusa is a common fern of North America, commonly known as the lowland bladderfern, lowland brittle fern or lowland fragile fern.
Cystopteris bulbifera, with the common name bulblet fern, bulblet bladderfern, or bulblet fragile fern is a fern in the family Cystopteridaceae.
Cystopteris tenuis is sometimes known as Mackay's bladder fern or Mackay's fragile fern. It was long considered to be a part of the superspecies for fragile ferns, as Cystopteris fragilis(L.) Bernh. var. mackayiLawson.
Bommeria is a genus of small pteridaceous rock ferns, native to the New World. Genetic analysis has shown it to be a stem offshoot clade of all the cheilanthoid ferns, except for Doryopteris, which is an even more basal ("primitive") stem offshoot. for years, it was assumed that this genus was closely allied with Hemionitis, but genetic analysis has shown that genus to be a more advanced genus evolutionarily. The same set of analysis has shown many presumed genera within this family to be paraphyletic, but the small genus Bommeria appears to be monophyletic, or a natural genus.
Cystopteris montana, previously classified as Athyrium montanum, is a species of fern known by the common name mountain bladderfern. It occurs throughout the high latitudes of the Northern Hemisphere, in Eurasia, Greenland, and Alaska, and throughout Canada. It is also present in the higher elevations in Colorado farther south.
Physematium neomexicanum, the New Mexican cliff fern, is a fern species native to the Southwestern United States and northern Mexico.
Myriopteris alabamensis, the Alabama lip fern, is a moderately-sized fern of the United States and Mexico, a member of the family Pteridaceae. Unlike many members of its genus, its leaves have a few hairs on upper and lower surfaces, or lack them entirely. One of the cheilanthoid ferns, it was usually classified in the genus Cheilanthes as Cheilanthes alabamensis until 2013, when the genus Myriopteris was again recognized as separate from Cheilanthes. It typically grows in shade on limestone outcrops.
Myriopteris aemula, the Texas lip fern or rival lip fern, is a moderately-sized fern of Texas and Mexico, a member of the family Pteridaceae. Unlike many members of its genus, its leaves have a few hairs on upper and lower surfaces, or lack them entirely. One of the cheilanthoid ferns, it was usually classified in the genus Cheilanthes as Cheilanthes aemula until 2013, when the genus Myriopteris was again recognized as separate from Cheilanthes. It typically grows on limestone rock.
Argyrochosma peninsularis is a fern endemic to Baja California Sur. It grows in dry, rocky places. First described as a species in 1939, it was transferred to the new genus Argyrochosma in 1987, recognizing their distinctness from the "cloak ferns". A dusting of powdery material and the presence of occasional scales on the central axis of its leaves help distinguish it from related species.
Argyrochosma pallens is a fern endemic to Mexico. It has narrow, divided leaves with brown axes; the leaves are dusted with white powder above and coated in it below. First described as a species in 1956, it was transferred to the new genus Argyrochosma in 1987, recognizing their distinctness from the "cloak ferns".
Argyrochosma fendleri, Fendler's false cloak fern, is a fern known from the western United States and northwestern Mexico. It grows in rocky habitats, and is distinguished from other members of the genus by its zig-zag leaf axes. Like many species in the genus, it bears white powder on the underside of its leaves. First described as a species in 1851, it was transferred to the new genus Argyrochosma in 1987, recognizing their distinctness from the "cloak ferns".
Argyrochosma limitanea, the southwestern false cloak fern, is a species of fern native to the southwestern United States and Sonora, Mexico. It grows on calcareous rocks, and has small, finely-divided leaves with a leathery texture, dark axes connecting the leaf segments, and a heavy coating of white powder on the undersurface. It reproduces apogamously; two subspecies are recognized, which may have originated independently through the hybridization of other taxa not yet discovered. First described as a species in 1919, it was transferred to the new genus Argyrochosma in 1987, recognizing their distinctness from the "cloak ferns".
Myriopteris rawsonii, formerly known as Cheilanthes rawsonii, is a perennial fern native to Namaqualand in Southern Africa. Like many other cheilanthoid ferns, it is adapted to dry conditions, bearing a thick layer of pale hairs on the underside of its pinnate-pinnatifid leaves. It is the only African representative of its clade of cheilanthoids, the otherwise American genus Myriopteris. It spends much of the year in a dried-out, dormant state, rehydrating and putting out new growth during winter rains. Its name honors the botanist and civil servant Sir Rawson W. Rawson.
Gymnocarpium disjunctum is a species of fern in the family Cystopteridaceae, commonly known as Pacific oak fern, western oak fern, or Pacific oakfern.
Myriopteris scabra, commonly known as rough lipfern, is a species of cheilanthoid fern native to Mexico and the south-western United States (Texas).
Myriopteris wrightii, formerly known as Cheilanthes wrightii, is a species of cheilanthoid fern with the common name Wright's lipfern. It is native to the southwestern United States and northern Mexico.
Cystopteris laurentiana, commonly called Laurentian bladderfern or St. Lawrence bladderfern, is a species of fern in the family Cystopteridaceae. It is native to eastern North America, primarily in the Great Lakes-St. Lawrence lowlands, but there are also scattered occurrences in New England and Atlantic Canada. It grows on cliffs composed of calcareous rocks, such as limestone, dolostone and diabase.
Astrolepis integerrima is a fern in the family Pteridaceae known by the common names hybrid cloakfern and southwest cloakfern. Though widespread through much of northern Mexico and parts of the southwestern and south central United States, it is nowhere common. It is only found on calcareous rocks, cliffs, and canyons in the deserts and similarly dry habitats. Astrolepis integerrima is an allotriploid, a type of hybrid containing all or almost all the chromosones of two different species.
