Dolichophonus | |
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†Dolichophonus loudonensis Laurie 1899. [1] |
Dolichophonus is an extinct genus of scorpions known from the Silurian-age Gutterford Burn Eurypterid bed, in the Pentland Hills in Scotland. It is currently considered the world's oldest known scorpion, as the older Parioscorpio named in 2020 has been found to be an unrelated arthropod. lt is sometimes referred to Palaeophonus . [2]
The specimen of Dolichophonus was collected by Mr Hardie of Bavelaw Castle. [1] When he died, his collection was given to the Edinburgh Museum of Science and Art (now National Museum of Scotland), where it was then studied by Malcolm Laurie. He named seven new species of eurypterids from the collection, along with this scorpion, which he named.
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event 251.9 million years ago.
Eurypterus is an extinct genus of eurypterid, a group of organisms commonly called "sea scorpions". The genus lived during the Silurian period, from around 432 to 418 million years ago. Eurypterus is by far the most well-studied and well-known eurypterid. Eurypterus fossil specimens probably represent more than 95% of all known eurypterid specimens.
Hibbertopterus is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of Hibbertopterus have been discovered in deposits ranging from the Devonian period in Belgium, Scotland and the United States to the Carboniferous period in Scotland, Ireland, the Czech Republic and South Africa. The type species, H. scouleri, was first named as a species of the significantly different Eurypterus by Samuel Hibbert in 1836. The generic name Hibbertopterus, coined more than a century later, combines his name and the Greek word πτερόν (pteron) meaning "wing".
Palaeophonus is the oldest known genus of scorpions.
Slimonia is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Slimonia have been discovered in deposits of Silurian age in South America and Europe. Classified as part of the family Slimonidae alongside the related Salteropterus, the genus contains three valid species, S. acuminata from Lesmahagow, Scotland, S. boliviana from Cochabamba, Bolivia and S. dubia from the Pentland Hills of Scotland and one dubious species, S. stylops, from Herefordshire, England. The generic name is derived from and honors Robert Slimon, a fossil collector and surgeon from Lesmahagow.
Carcinosoma is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Carcinosoma are restricted to deposits of late Silurian age. Classified as part of the family Carcinosomatidae, which the genus lends its name to, Carcinosoma contains seven species from North America and Great Britain.
Chasmataspidids, sometime referred to as chasmataspids, are a group of extinct chelicerate arthropods that form the order Chasmataspidida. Chasmataspidids are probably related to horseshoe crabs (Xiphosura) and/or sea scorpions (Eurypterida), with more recent studies suggest that they form a clade (Dekatriata) with Eurypterida and Arachnida. Chasmataspidids are known sporadically in the fossil record through to the mid-Devonian, with possible evidence suggesting that they were also present during the late Cambrian. Chasmataspidids are most easily recognised by having an opisthosoma divided into a wide forepart (preabdomen) and a narrow hindpart (postabdomen) each comprising 4 and 9 segments respectively. There is some debate about whether they form a natural group.
Drepanopterus is an extinct genus of eurypterid and the only member of the family Drepanopteridae within the Mycteropoidea superfamily. There are currently three species assigned to the genus. The genus has historically included more species, with nine species associated with the genus Drepanopterus, however five of these have since been proven to be synonyms of pre-existing species, assigned to their own genera, or found to be based on insubstantial fossil data. The holotype of one species proved to be a lithic clast.
Campylocephalus is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of Campylocephalus have been discovered in deposits ranging from the Carboniferous period in the Czech Republic to the Permian period of Russia. The generic name is composed of the Greek words καμπύλος (kampýlos), meaning "curved", and κεφαλή (kephalē), meaning "head".
Onychopterella is a genus of predatory eurypterid, an extinct group of aquatic arthropods. Fossils of Onychopterella have been discovered in deposits from the Late Ordovician to the Late Silurian. The genus contains three species: O. kokomoensis, the type species, from the Early Pridoli epoch of Indiana; O. pumilus, from the Early Llandovery epoch of Illinois, both from the United States; and O. augusti, from the Late Hirnantian to Early Rhuddanian stages of South Africa.
Kiaeropterus is a genus of prehistoric eurypterid classified as part of the Rhenopteridae family. Two species, both from the Silurian period, are known; K. cyclophthalmus from Scotland and K. ruedemanni from Norway.
Parastylonurus is a genus of prehistoric eurypterid. It is classified within the Parastylonuridae family and contains three species, P. hendersoni and P. ornatus from the Silurian of Scotland and P. sigmoidalis from the Silurian of England.
Lanarkopterus is a genus of prehistoric eurypterid with one recognised species, Lanarkopterus dolichoschelus. Lanarkopterus was long seen as a species of the closely related Mixopterus, though more complete specimens discovered in the 1960s determined that it differed in several aspects, enough to warrant a separate genus.
Hastimima is a genus of prehistoric eurypterid of the family Mycteroptidae. It contains one valid species, Hastimima whitei, recovered from the Permian Hermit Shale of Arizona, and one dubious species, H. sewardi, from the Devonian of South Africa. Though clearly a mycteropoid eurypterid, the material referred to H. sewardi is so fragmentary that it is questionable if it possesses any diagnostic value at all.
Bembicosoma is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Bembicosoma was regarded as part of the clade Planaterga. Fossils of the single and type species, B. pomphicus, have been discovered in deposits of the Silurian period in the Pentland Hills, Scotland. Bembicosoma had been tentatively assigned as an eurypterid before its synziphosurine affinities revealed.
Hibbertopteridae is a family of eurypterids, an extinct group of aquatic arthropods. They were members of the superfamily Mycteropoidea. Hibbertopterids were large, broad and heavy animals unlike virtually every other group of eurypterids, which are commonly streamlined and lightweight. Their bizarre morphology is so unusual that they in the past have been thought to represent an entirely distinct order of chelicerates. Fossils of the family first appear in deposits of Middle Devonian age and the last known fossils representing hibbertopterids are known from deposits of Late Permian age. The hibbertopterids represent the last known living eurypterids, going extinct during the Permian–Triassic extinction event or shortly before.
Vernonopterus is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of Vernonopterus have been discovered in deposits of the Carboniferous period in Scotland. The name of the genus derives from the location where the only known fossil has been discovered, Mount Vernon near Airdrie in Lanarkshire, Scotland. A single species of Vernonopterus is recognized, V. minutisculptus, based on fragmentary fossilized tergites, segments on the upper side of the abdomen. The species name minutisculptus refers to the ornamentation of scales that covers the entirety of the preserved parts of the eurypterid.
Eusarcana is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Eusarcana have been discovered in deposits ranging in age from the Early Silurian to the Early Devonian. Classified as part of the family Carcinosomatidae, the genus contains three species, E. acrocephalus, E. obesus and E. scorpionis, from the Silurian-Devonian of Scotland, the Czech Republic and the United States respectively.
This timeline of eurypterid research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of eurypterids, a group of extinct aquatic arthropods closely related to modern arachnids and horseshoe crabs that lived during the Paleozoic Era.