The fiddler crab or calling crab may be any of more than one hundred species of semiterrestrial marine crabs in the family Ocypodidae, well known for their sexually dimorphic claws; the males' major claw is much larger than the minor claw, while the females' claws are both the same size. A smaller number of ghost crab and mangrove crab species are also found in the family Ocypodidae. This entire group is composed of small crabs, the largest being slightly over two inches (5 cm) across. Fiddler crabs are found along sea beaches and brackish intertidal mud flats, lagoons, swamps, and various other types of brackish or salt-water wetlands.
Xanthidae is a family of crabs known as gorilla crabs, mud crabs, pebble crabs or rubble crabs. Xanthid crabs are often brightly coloured and are highly poisonous, containing toxins which are not destroyed by cooking and for which no antidote is known. The toxins are similar to the tetrodotoxin and saxitoxin produced by puffer fish, and may be produced by bacteria in the genus Vibrio living in symbiosis with the crabs, mostly V. alginolyticus and V. parahaemolyticus.
Hymenosomatidae is a family of crabs with some 110 described species in nearly 20 genera. The following genera are placed in the Hymenosomatidae:
Sand bubbler crabs are crabs of the genera Scopimera and Dotilla in the family Dotillidae. They are small crabs that live on sandy beaches in the tropical Indo-Pacific. They feed by filtering sand through their mouthparts, leaving behind balls of sand that are disintegrated by the incoming high tide.
Macrophthalmus is a genus of crabs which are widespread across the Indo-Pacific. It contains the following species :
Discoplax is a genus of terrestrial crabs. It is very closely related to the genus Cardisoma.
Pseudothelphusidae is a family of freshwater crabs found chiefly in mountain streams in the Neotropics. They are believed to have originated in the Greater Antilles and then crossed to Central America via a Pliocene land bridge.
The Sesarmidae are a family of crabs, previously included in the Grapsidae by many authors. Several species, namely in Geosesarma, Metopaulias, and Sesarma, are true terrestrial crabs. They do not need to return to the sea even for breeding.
Geosesarma is genus of small freshwater or terrestrial crabs, typically less than 10 mm (0.4 in) across the carapace. They live and reproduce on land with the larval stages inside the egg. They are found from India, through Southeast Asia, to the Solomon Islands and Hawaii.
The Varunidae are a family of thoracotrematan crabs. The delimitation of this family, part of the taxonomically confusing Grapsoidea, is undergoing revision. For a long time, they were placed at the rank of subfamily in the Grapsidae, but they appear to be closest to Macropthalmus and the Mictyridae, which are usually placed in the Ocypodoidea. It may thus be better to merge the latter superfamily with the Grapsoidea, retaining the latter name as it is older.
Pilumnoidea is a superfamily of crabs, whose members were previously included in the Xanthoidea. The three families are unified by the free articulation of all the segments of the male crab's abdomen and by the form of the gonopods. The earliest fossils assigned to this group are of Eocene age.
Hemigrapsus is a genus of varunid crabs comprising thirteen species native almost exclusively in the Pacific Ocean, but two have been introduced to the North Atlantic region.
Karstarma is a genus of karst-dwelling crabs formerly included in Sesarmoides.
Leucosiidae is a family of crabs containing three subfamilies and a number of genera incertae sedis:
Geograpsus is a genus of crabs in the family Grapsidae, containing four extant species, and one extinct species:
Glyptoxanthus is a genus of crabs in the family Xanthidae, containing eight species. It was originally erected by Alphonse Milne-Edwards in 1879 for six species previously placed in the genus Actaea and elsewhere. Although previously included in subfamily Euxanthinae, the genus has a quite distinct morphology from other genera in that group, and was placed in 2011 in the new, monotypic subfamily, Glyptoxanthinae by Jose Christopher Mendoza and Danièle Guinot.
Ebalia is a genus of crab in the family Leucosiidae.
Paragrapsus is a genus of crabs from South Eastern Australia, containing the following species:
Limnopilos naiyanetri, commonly referred to as the Thai micro crab or pill-box crab, is a freshwater hymenosomatid crab endemic to Thailand. Its presence has only been confirmed in the Tha Chin River. The species was described in 1991 and represents the type species of Limnopilos. The Thai micro crab was first introduced to the aquarium hobby in 2008 when it was imported to Germany by the tropical fish importer Aquarium Glaser GmbH, and has slowly grown in popularity with aquarium hobbyists. It remains a relatively rare species on the market and detailed information on the husbandry of this species is scarce.
Limnopilos is a genus of small hymenosomatid crabs endemic to Southeast Asia. The genus was described by Christina Chuang and Peter Ng in 1991, who identified the new species Limnopilos naiyanetri and distinguished it from the closely related genus Hymenicoides. Its true taxonomic classification was debated for several years, but in 2007 the discovery of a new species of crab in this genus solidified the distinction between Limnopilos and Hymenicoides. Many aspects of the genus Limnopilos are still poorly understood. Their ecology and natural history have not been studied in detail, and their reproductive cycle remains mysterious.
