Electrostephanus Temporal range: | |
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Electrostephanus petiolatus neotype male | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Family: | Stephanidae |
Subfamily: | † Electrostephaninae |
Genus: | † Electrostephanus Brues, 1933 |
Subgenera | |
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Electrostephanus is an extinct genus of crown wasp in the hymenopteran family Stephanidae, and is the only genus placed in the subfamily Electrostephaninae. The genus contains four described species, E. brevicornis, E. neovenatus, E. janzeni, and E. petiolatus, placed in two subgenera E. (Electrostephanus) and E. (Electrostephanodes). Electrostephanus is known from several middle Eocene fossils which have been found in Europe. [1] [2]
Electrostephanus is known from a group of fossils preserved as an inclusions in a transparent chunks of Baltic amber and is named after the Greek term for amber, ἤλεκτρον (elektron). Baltic amber is approximately forty six million years old, having been deposited during Lutetian stage of the Middle Eocene. There is debate as to what plant family the amber was produced by, with evidence supporting relatives of either Agathis or Pseudolarix trees. [3] The genus was originally described by paleoentomologist Charles Thomas Brues in 1933 with Brues designating E. petiolatus as the type species, and including two other species, E. tridentatus and E. brevicornis. A fourth species, "E." sulcatus and fifth, E. neovenatus, were described by Alexandre Aguiar and Jens-Wilhelm Janzen 1999, and a sixth species. E. janzeni, was described in 2005. [1] The original type specimens for E. petiolatus, "E." tridentatus and "E." brevicornis were part of the Albertus Universität, Königsberg collection of ambers, and all were possibly lost to fire during the bombing of Kaliningrad in World War II. [1]
Based on the structure of the petiole and abdomen in "E.". sulcatus and the suggestion that E. petiolatus had the same morphology, Electrostephanus was designated a junior synonym of Denaeostephanus in 2004. However, this move was reversed with the description of an additional male E. petiolatus specimen in 2008 by Michael S. Engel and Jaime Ortega-Blanco, who designated the new specimen the neotype for E. petiolatus. Engel and Ortega-Blanco transferred the species back to Electrostephanus along with E. brevicornis, E. neovenatus, and E. janzeni, but retaining D. tridentatus in Denaeostephanus. Based on the abdomen morphology Engel and Ortega-Blanco further split the genus into two distinct subgenera, E. (Electrostephanus) containing the type species E. petiolatus, and E. (Electrostephanodes) containing the remaining three species. They noted that Electrostephanodes may merit elevation to full genus status, but the lack of additional known fossil specimens to consult meant they opted for a conservative subgenus placement. [1]
The Electrostephanus petiolus male adult has an elongated body which is 9.84 mm (0.387 in) long with an overall coloration ranging from black to dark brown and having scattered setae. The head capsule is generally spherical with rounded compound eyes on the lateral surface. The typical "crown" is a group of five tubercles places in front of four small ridges running crossways over the head capsule. The antennae are composed of 22 flagellomeres and scape, attaching to the head near the mid-line of the compound eyes. The wings are hyaline in coloration, with a dark brown pterostigma that is parallel sided and the estimated fore-wing length is 5.04 mm (0.198 in). The metasoma is narrow and forms a petiole. [1]
The Stephanidae, sometimes called crown wasps, are a family of parasitoid wasps. They are the only living members of the superfamily Stephanoidea. Stephanidae has at least 345 living species in 11 genera. The family is considered cosmopolitan in distribution, with the highest species concentrations in subtropical and moderate climate zones. Stephanidae also contain four extinct genera described from both compression fossils and inclusions in amber.
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Deinodryinus velteni is an extinct species of Deinodryinus in the wasp family Dryinidae. The species is known solely from an Eocene fossil found in the Baltic region.
Astreptolabis is an extinct genus of earwig in the Dermaptera family Pygidicranidae known from a group of Cretaceous fossils found in Myanmar. The genus contains two described species, Astreptolabis ethirosomatia and Astreptolabis laevis and is the sole member of the subfamily Astreptolabidinae.
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Prostylotermes is an extinct genus of termite in the isopteran family Stylotermitidae known from two Eocene fossils found in India. The genus contains a single described species, Prostylotermes kamboja.
Termitaradus avitinquilinus is an extinct species of termite bug in the family Termitaphididae known from several possibly Miocene fossils found in the Dominican Republic. T. avitinquilinus is the first species in the genus Termitaradus to have been described from fossils found in Dominican amber and is one of four species from New World amber, the others being Termitaradus protera, Termitaradus dominicanus and Termitaradus mitnicki.
Yantaromyrmex is an extinct genus of ants first described in 2013. Members of this genus are in the subfamily Dolichoderinae of the family Formicidae, known from Middle Eocene to Early Oligocene fossils found in Europe. The genus currently contains five described species, Y. constrictus, Y. geinitzi, Y. intermedius, Y. mayrianum and Y. samlandicus. The first specimens were collected in 1868 and studied by Austrian entomologist Gustav Mayr, who originally placed the fossils in other ant genera until the fossils were reviewed and subsequently placed into their own genus. These ants are small, measuring from 4 to 6 mm in length and can be characterized by their trapezoidal shaped head-capsules and oval compound eyes that are located slightly to the rear of the capsules midpoint, with no known ocelli present.
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Embolemidae is a family of small solitary parasitoid wasps with around 70 species in 2 genera distributed around the world. The few species whose biology is known are parasites on planthopper nymphs of the families Achilidae and Cixiidae. There is debate regarding the status of the genus named Ampulicomorpha by Ashmead in 1893, generally considered now to be a junior synonym of Embolemus (e.g.,), though some authorities dispute this (e.g.,)
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