Pteranodon is a genus of pterosaur that included some of the largest known flying reptiles, with P. longiceps having a wingspan of over 6 m (20 ft). They lived during the late Cretaceous geological period of North America in present-day Kansas, Nebraska, Wyoming, South Dakota and Alabama. More fossil specimens of Pteranodon have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway.
The Western Interior Seaway was a large inland sea that split the continent of North America into two landmasses for 34 million years. The ancient sea, which existed from the early Late Cretaceous to the earliest Paleocene, connected the Gulf of Mexico to the Arctic Ocean. The two land masses it created were Laramidia to the west and Appalachia to the east. At its largest extent, it was 2,500 feet (760 m) deep, 600 miles (970 km) wide and over 2,000 miles (3,200 km) long.
Archelon is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring 4.6 m (15 ft) from head to tail and 2.2–3.2 t in body mass. It is known only from the Pierre Shale and has one species, A. ischyros. In the past, the genus also contained A. marshii and A. copei, though these have been reassigned to Protostega and Kansastega, respectively. The genus was named in 1895 by American paleontologist George Reber Wieland based on a skeleton from South Dakota, who placed it into the extinct family Protostegidae. The leatherback sea turtle was once thought to be its closest living relative, but now, Protostegidae is thought to be a completely separate lineage from any living sea turtle.
Cretoxyrhina is an extinct genus of large mackerel shark that lived about 107 to 73 million years ago during the late Albian to late Campanian of the Late Cretaceous. The type species, C. mantelli, is more commonly referred to as the Ginsu shark, first popularized in reference to the Ginsu knife, as its theoretical feeding mechanism is often compared with the "slicing and dicing" when one uses the knife. Cretoxyrhina is traditionally classified as the likely sole member of the family Cretoxyrhinidae but other taxonomic placements have been proposed, such as within the Alopiidae and Lamnidae.
Ichthyornis is an extinct genus of toothy seabird-like ornithuran from the late Cretaceous period of North America. Its fossil remains are known from the chalks of Alberta, Alabama, Kansas, New Mexico, Saskatchewan, and Texas, in strata that were laid down in the Western Interior Seaway during the Turonian through Campanian ages, about 95–83.5 million years ago. Ichthyornis is a common component of the Niobrara Formation fauna, and numerous specimens have been found.
Enchodus is an extinct genus of aulopiform ray-finned fish related to lancetfish and lizardfish. Species of Enchodus flourished during the Late Cretaceous, and there is some evidence that they may have survived to the Paleocene or Eocene; however, this may just represent reworked Cretaceous material.
Xiphactinus, colloquially referred to as the X-fish, is an extinct genus of large predatory marine bony fish that lived during the late Albian to the late Maastrichtian. The genus grew up to 5–6 metres (16–20 ft) in length, and superficially resembled a gargantuan, fanged tarpon.
Protosphyraena is a fossil genus of swordfish-like marine fish, that thrived worldwide during the Upper Cretaceous Period (Coniacian-Maastrichtian). Though fossil remains of this taxon have been found in both Europe and Asia, it is perhaps best known from the Smoky Hill Member of the Niobrara Chalk Formation of Kansas. Protosphyraena was a large fish, averaging 2–3 metres in length. Protosphyraena shared the Cretaceous oceans with aquatic reptiles, such as mosasaurs and plesiosaurs, as well as with many other species of extinct predatory fish. The name Protosphyraena is a combination of the Greek word protos ("early") plus Sphyraena, the genus name for barracuda, as paleontologists initially mistook Protosphyraena for an ancestral barracuda. Recent research shows that the genus Protosphyraena is not at all related to the true swordfish-family Xiphiidae, but belongs to the extinct family Pachycormidae.
Nyctosaurus is a genus of nyctosaurid pterosaur from the Late Cretaceous period of what is now the Niobrara Formation of the mid-western United States, which, during the time Nyctosaurus was alive, was covered in an extensive shallow sea. Some remains belonging to a possible Nyctosaurus species called "N." lamegoi have been found in Brazil, though it likely belongs to a different genus Simurghia. The genus Nyctosaurus has had numerous species referred to it, though how many of these may actually be valid requires further study. At least one species possessed an extraordinarily large antler-like cranial crest.
Protostega is an extinct genus of sea turtle containing a single species, Protostega gigas. Its fossil remains have been found in the Smoky Hill Chalk formation of western Kansas, time-equivalent beds of the Mooreville Chalk Formation of Alabama and Campanian beds of the Rybushka Formation. Fossil specimens of this species were first collected in 1871, and named by Edward Drinker Cope in 1872. With a total length of 3.9 metres (13 ft), it is the second-largest sea turtle that ever lived, second only to the giant Archelon, and one of the three largest turtle of all time along Archelon and Gigantatypus.
Dolichorhynchops is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing the species D. osborni and D. herschelensis, with two previous species having been assigned to new genera. Definitive specimens of D. osborni have been found in the late Coniacian to early Campanian rocks, while those of D. herschelensis have been found in the late Campanian to early Maastrichtian rocks. Dolichorhynchops was a prehistoric marine reptile measuring around 3 metres (9.8 ft) long. Its Greek generic name means "long-nosed face".
Tusoteuthis is an extinct genus of large enchoteuthine cephalopod that lived during the Cretaceous. Although often called a squid, it is now thought to be more closely related to modern octopuses. Although many specimens from Western Interior Seaway of North America were described, at 2019, only one, poorly preserved specimen from Kansas is recognized as Tusoteuthis. One species, T. longa, is traditionally recognized. In 2019, due to poor preservation of holotype specimen, Tusoteuthis was considered likely to be a nomen dubium, and it was proposed that later described species are better to included in genus Enchoteuthis instead.
Styxosaurus is a genus of plesiosaur of the family Elasmosauridae. Styxosaurus lived during the Campanian age of the Cretaceous period. Three species are known: S. snowii, S. browni, and S. rezaci.
The Pierre Shale is a geologic formation or series in the Upper Cretaceous which occurs east of the Rocky Mountains in the Great Plains, from Pembina Valley in Canada to New Mexico.
The Niobrara Formation, also called the Niobrara Chalk, is a geologic formation in North America that was deposited between 87 and 82 million years ago during the Coniacian, Santonian, and Campanian stages of the Late Cretaceous. It is composed of two structural units, the Smoky Hill Chalk Member overlying the Fort Hays Limestone Member. The chalk formed from the accumulation of coccoliths from microorganisms living in what was once the Western Interior Seaway, an inland sea that divided the continent of North America during much of the Cretaceous. It underlies much of the Great Plains of the US and Canada. Evidence of vertebrate life is common throughout the formation and includes specimens of plesiosaurs, mosasaurs, pterosaurs, and several primitive aquatic birds. The type locality for the Niobrara Chalk is the Niobrara River in Knox County in northeastern Nebraska. The formation gives its name to the Niobrara cycle of the Western Interior Seaway.
During the time of the deposition of the Niobrara Chalk, much life inhabited the seas of the Western Interior Seaway. By this time in the Late Cretaceous many new lifeforms appeared such as mosasaurs, which were to be some of the last of the aquatic lifeforms to evolve before the end of the Mesozoic. Life of the Niobrara Chalk is comparable to that of the Dakota Formation, although the Dakota Formation, which was deposited during the Cenomanian, predates the chalk by about 10 million years.
Cretodus is an extinct genus of large mackerel sharks belonging to the proposed family Pseudoscapanorhynchidae. Cretodus lived during the Late Cretaceous, ranging from the Cenomanian to the Coniacian. The genus is well-known from strata deposited in the Western Interior Seaway, and from the Late Cretaceous of Europe, Africa, and possibly Asia. Cretodus is primarily represented in the fossil record by isolated teeth and vertebral centra, though a couple of associated dentitions and vertebral columns have been found.
Marion Charles Bonner (1911–1992), based for most of his life in Leoti, Kansas, was an American field paleontologist who discovered and collected hundreds of fossils, primarily from the Niobrara Cretaceous Smoky Hill chalk outcroppings in Logan, Scott, and Gove counties of western Kansas. Largely self-taught, he frequently collaborated with museum paleontologists, including George F. Sternberg, at Fort Hays State University in Hays, Kansas, and Shelton P. Applegate, at the Los Angeles County Museum of Natural History.
Platylithophycus is an extinct genus of elasmobranchs that lived during the Late Cretaceous. It is known from a single specimen from the Niobrara Formation of Kansas, United States. It was originally identified as the fronds of a codiacean alga, then later as the cuttlebone of a cuttlefish. It was most recently reidentified as the gill arches and rakers of an elasmobranch of uncertain affinities. It might have been a filter feeding mackerel shark related to Aquilolamna.
Martinectes is an extinct genus of polycotylid plesiosaur from the Late Cretaceous Sharon Springs Formation of the United States. The genus contains a single species M. bonneri, known from multiple skeletons and skulls. Martinectes was historically considered to represent a species of the genus Trinacromerum and later Dolichorhynchops before it was moved to its own genus. It was a large polycotylid measuring around 6–7 metres (20–23 ft) long.
