Pteranodon ; from Ancient Greek πτερόν and ἀνόδων is a genus of pterosaur that included some of the largest known flying reptiles, with P. longiceps having a wingspan of over 6 m (20 ft). They lived during the late Cretaceous geological period of North America in present-day Kansas, Nebraska, Wyoming, South Dakota and Alabama. More fossil specimens of Pteranodon have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway.
The Western Interior Seaway was a large inland sea that split the continent of North America into two landmasses. The ancient sea, which existed from the early Late Cretaceous to the earliest Paleocene, connected the Gulf of Mexico, through the United States and Canada, to the Arctic Ocean. The two land masses it created were Laramidia to the west and Appalachia to the east. At its largest extent, it was 2,500 feet (760 m) deep, 600 miles (970 km) wide and over 2,000 miles (3,200 km) long.
Archelon is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring 4.6 m (15 ft) from head to tail and 2.2–3.2 t in body mass. It is known only from the Pierre Shale and has one species, A. ischyros. In the past, the genus also contained A. marshii and A. copei, though these have been reassigned to Protostega and Microstega, respectively. The genus was named in 1895 by American paleontologist George Reber Wieland based on a skeleton from South Dakota, who placed it into the extinct family Protostegidae. The leatherback sea turtle was once thought to be its closest living relative, but now, Protostegidae is thought to be a completely separate lineage from any living sea turtle.
Ichthyornis is an extinct genus of toothy seabird-like ornithuran from the late Cretaceous period of North America. Its fossil remains are known from the chalks of Alberta, Alabama, Kansas, New Mexico, Saskatchewan, and Texas, in strata that were laid down in the Western Interior Seaway during the Turonian through Campanian ages, about 95–83.5 million years ago. Ichthyornis is a common component of the Niobrara Formation fauna, and numerous specimens have been found.
Enchodus is an extinct genus of aulopiform ray-finned fish related to lancetfish and lizardfish. Species of Enchodus flourished during the Late Cretaceous, and survived the Cretaceous–Paleogene extinction event, persisting into the late Eocene.
Niobrarasaurus is an extinct genus of nodosaurid ankylosaur which lived during the Cretaceous 87 to 82 million years ago. Its fossils were found in the Smoky Hill Chalk Member of the Niobrara Formation, in western Kansas, which would have been near the middle of Western Interior Sea during the Late Cretaceous. It was a nodosaurid, an ankylosaur without a clubbed tail. It was closely related to Nodosaurus.
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The formation stretches over portions of Montana, North Dakota, South Dakota, and Wyoming. In Montana, the Hell Creek Formation overlies the Fox Hills Formation. The site of Pompeys Pillar National Monument is a small isolated section of the Hell Creek Formation. In 1966, the Hell Creek Fossil Area was designated as a National Natural Landmark by the National Park Service.
Nyctosaurus is a genus of nyctosaurid pterosaur from the Late Cretaceous period of what is now the Niobrara Formation of the mid-western United States, which, during the time Nyctosaurus was alive, was covered in an extensive shallow sea. Some remains belonging to a possible Nyctosaurus species called N.lamegoi have been found in Brazil, making Nyctosaurus more diverse. The genus Nyctosaurus has had numerous species referred to it, though how many of these may actually be valid requires further study. At least one species possessed an extraordinarily large antler-like cranial crest.
Protostega is an extinct genus of sea turtle containing a single species, Protostega gigas. Its fossil remains have been found in the Smoky Hill Chalk formation of western Kansas, time-equivalent beds of the Mooreville Chalk Formation of Alabama and Campanian beds of the Rybushka Formation. Fossil specimens of this species were first collected in 1871, and named by Edward Drinker Cope in 1872. With a total length of 3.9 metres (13 ft), it is the second-largest sea turtle that ever lived, second only to the giant Archelon, and one of the three largest turtle of all time along Archelon and Gigantatypus.
Dolichorhynchops is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing three species, D. osborni, D. bonneri and D. tropicensis, as well as a questionably referred fourth species, D. herschelensis. Dolichorhynchops was a prehistoric marine reptile, but at least one species, D. tropicensis, likely entered rivers to collect gastroliths. Its Greek generic name means "long-nosed face". While typically measuring about 3 metres (9.8 ft) in length, the largest specimens of D. osborni and D. bonneri are estimated to have a total body length more than approximately 4.29 metres (14.1 ft) and 5.09 metres (16.7 ft), respectively.
Tusoteuthis is an extinct genus of large enchoteuthine cephalopod that lived during the Cretaceous. Although often called a squid, it is now thought to be more closely related to modern octopuses. Examination of gladius remains has yielded an estimated mantle length close to or equal to that of the modern giant squid. Fossil remains have been found in parts of the one-time Western Interior Seaway of North America, including Late Cretaceous rocks in Kansas, Colorado, Wyoming, South Dakota, North Dakota, and Manitoba Province. One species, T. longa, is traditionally recognized. In 2019, due to poor preservation of holotype specimen, genus Tusoteuthis is likely to be invalid, and it is considered that later described species are better to include in genus Enchoteuthis instead.
Styxosaurus is a genus of plesiosaur of the family Elasmosauridae. Styxosaurus lived during the Campanian age of the Cretaceous period. Two species are known: S. snowii and S. browni.
Ptychodus is a genus of extinct durophagous (shell-crushing) sharks from the Late Cretaceous. Fossils of Ptychodus teeth are found in many Late Cretaceous marine sediments. There are many species among the Ptychodus that have been uncovered on all the continents around the globe. Such species are Ptychodus mortoni, P. decurrens, P. marginalis, P. mammillaris, P. rugosus and P. latissimus to name a few. They died out approximately 85 million years ago. A large number of remains have been found in the former Western Interior Seaway. A 2016 publication found that Ptychodus are likely true sharks belonging to Selachimorpha, rather than hybodonts or batoids as previously thought. Their life history coincides with the typical life of many other large sharks: they lived relatively long lives and were slow growing and produced large offspring and small litters.
Stratodus is a genus of giant prehistoric aulopiform fish found in Cretaceous-aged marine strata of Kansas, Alabama, Morocco, Israel, and Niger, South Dakota, Jordan. It has also been found in the Tamaguélelt Formation of Mali, dating to the Lower Eocene, indicating that Stratodus survived the Cretaceous-Paleogene extinction event. This sleek fish has an upper jaw filled with multiple rows of tiny teeth and was the largest aulopiform, reaching 5 meters in length.
The Pierre Shale is a geologic formation or series in the Upper Cretaceous which occurs east of the Rocky Mountains in the Great Plains, from Pembina Valley in Canada to New Mexico.
Toxochelys is an extinct genus of marine turtle from the Late Cretaceous period. It is the most commonly found fossilized turtle species in the Smoky Hill Chalk, in western Kansas.
The Niobrara Formation, also called the Niobrara Chalk, is a geologic formation in North America that was deposited between 87 and 82 million years ago during the Coniacian, Santonian, and Campanian stages of the Late Cretaceous. It is composed of two structural units, the Smoky Hill Chalk Member overlying the Fort Hays Limestone Member. The chalk formed from the accumulation of coccoliths from microorganisms living in what was once the Western Interior Seaway, an inland sea that divided the continent of North America during much of the Cretaceous. It underlies much of the Great Plains of the US and Canada. Evidence of vertebrate life is common throughout the formation and includes specimens of plesiosaurs, mosasaurs, and pterosaurs as well as several primitive aquatic birds. The type locality for the Niobrara Chalk is the Niobrara River in Knox County in northeastern Nebraska. The formation gives its name to the Niobrara cycle of the Western Interior Seaway.
During the time of the deposition of the Niobrara Chalk, much life inhabited the seas of the Western Interior Seaway. By this time in the Late Cretaceous many new lifeforms appeared such as mosasaurs, which were to be some of the last of the aquatic lifeforms to evolve before the end of the Mesozoic. Life of the Niobrara Chalk is comparable to that of the Dakota Formation, although the Dakota Formation, which was deposited during the Cenomanian, predates the chalk by about 10 million years.
Paleontology in Iowa refers to paleontological research occurring within or conducted by people from the U.S. state of Iowa. The paleozoic fossil record of Iowa spans from the Cambrian to Mississippian. During the early Paleozoic Iowa was covered by a shallow sea that would later be home to creatures like brachiopods, bryozoans, cephalopods, corals, fishes, and trilobites. Later in the Paleozoic, this sea left the state, but a new one covered Iowa during the early Mesozoic. As this sea began to withdraw a new subtropical coastal plain environment which was home to duck-billed dinosaurs spread across the state. Later this plain was submerged by the rise of the Western Interior Seaway, where plesiosaurs lived. The early Cenozoic is missing from the local rock record, but during the Ice Age evidence indicates that glaciers entered the state, which was home to mammoths and mastodons.
Marion Charles Bonner (1911–1992), based for most of his life in Leoti, Kansas, was an American field paleontologist who discovered and collected hundreds of fossils, primarily from the Niobrara Cretaceous Smoky Hill chalk outcroppings in Logan, Scott, and Gove counties of western Kansas. Largely self-taught, he frequently collaborated with museum paleontologists, including George F. Sternberg, at Fort Hays State University in Hays, Kansas, and Shelton P. Applegate, at the Los Angeles County Museum of Natural History.
