Endoceras

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Endoceras
Temporal range: M-U Ordovician
Endoceras.JPG
Specimens in Milan
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Nautiloidea
Order: Endocerida
Family: Endoceratidae
Genus: Endoceras
Hall, 1847

Endoceras (Ancient Greek for "inner horn") is an extinct genus of large, straight shelled cephalopods that gives its name to the Nautiloid order Endocerida. The genus lived during the middle and upper Ordovician 485 to 419 million years ago. The cross section in the mature portion is slightly wider than high, but is narrower laterally in the young. Sutures are straight and transverse. Endoceras has a large siphuncle, located close to the ventral margin, composed of concave segments, especially in the young but which may be tubular in the adult stage. Endocones are simple, subcircular in cross section, and penetrated by a narrow tube which may contain diaphragms reminiscent of the Ellesmerocerid ancestor.

Specimen from Stevns, Denmark Endoceras.jpg
Specimen from Stevns, Denmark

Endoceras was named by Hall in 1847. Distribution is widespread, especially in North America and Europe; and fossils have been found in Australia. [1] Endoceras is similar to Cameroceras , the two may be synonymous, but differs from the genus Nanno in that the siphuncle in Nanno fills the entire apical portion of the shell while in Endoceras the siphuncle is ventral even there with septa formed at the onset.

Mature, full grown, Endoceras were most likely ambush predators that lay in wait on the sea floor, moving when necessary to gain the advantage. Younger individuals with compressed cross sections may have been more actively mobile.

Size

Life restoration of Endoceras giganteum Endoceras life restoration.png
Life restoration of Endoceras giganteum

A specimen of Endoceras giganteum at the Museum of Comparative Zoology measures 3 meters (9.8 feet) as preserved. [2] The most recent estimate puts its complete size at approximately 5.73 meters (18.8 feet). This would make it the largest cephalopod by length in the fossil record. [3] There is additionally an unconfirmed report of a 9.1-meter (30-foot) shell that was destroyed. [4]

Related Research Articles

<span class="mw-page-title-main">Nautiloid</span> Extant subclass of cephalopods

Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and species rich, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). No orthoconic and only a handful of coiled species, the nautiluses, survive to the present day.

<span class="mw-page-title-main">Endocerida</span> Fossil order of cephalopods

Endocerida is an extinct nautiloid order, a group of cephalopods from the Lower Paleozoic with cone-like deposits in their siphuncle. Endocerida was a diverse group of cephalopods that lived from the Early Ordovician possibly to the Late Silurian. Their shells were variable in form. Some were straight (orthoconic), others curved (cyrtoconic); some were long (longiconic), others short (breviconic). Some long-shelled forms like Endoceras attained shell lengths close to 6 metres (20 ft). The related Cameroceras is anecdotally reported to have reached lengths approaching 9 metres (30 ft), but these claims are problematic. The overwhelming majority of endocerids and nautiloids in general are much smaller, usually less than a meter long when fully grown.

Cameroceras is an extinct genus of endocerid cephalopod which lived in equatorial oceans during the entire Ordovician period. Like other endocerids, it was an orthocone, meaning that its shell was fairly straight and pointed. It was particularly abundant and widespread in the Late Ordovician, inhabiting the shallow tropical seas in and around Laurentia, Baltica and Siberia.

Suecoceras is an endoceratid that lived during the Middle Ordovician. It is characterised by a long, straight, slender shell with a slightly expanded tip that curves slightly downwards.

The Proterocameroceratidae were the first of the Endocerida. They began early in the Ordovician with Proendoceras or similar genus which had developed endocones, replacing the diaphragms of the ellesmerocerid ancestor.

Actinoceras is the principal and root genus of the Actinoceratidae, a major family in the Actinocerida, that lived during the Middle and Late Ordovician. It is an extinct genus of nautiloid cephalopod that thrived in the warm waters of the United States and England during the Paleozoic era.

Lambeoceras is a genus of large actinocerids with a convexly lenticular cross section from the Upper Ordovician of North America and the sole representative of the family Lambeoceratidae.

Armenoceras is a genus of actinocerid nautiloid cephalopods whose fossils ranged from the late Whiterockian Stage in the early Middle Ordovician, through the remainder of the period and on into the Upper Silurian. It is the type genus of the family Armenoceratidae.

Gonioceras is an extinct genus of actinocerid nautiloidean cephalopods typified by a broad, low shell; flattened ventrally, convexly rounded dorsally; top and bottom meeting at an acute angle along the sides. In most the shell is rather thin, especially along the lateral portion. The aperture is contracted. Sutures from broad ventral and dorsal lobes, more narrowly rounded ventro-lateral and dorso-lateral saddles, and sharp pointed lateral lobes; more complex than in later Lambeoceras. The siphuncle is typically subcentral but may be closer to the venter; armenocerid in form with short segments and very short brims and containing a straight endosiphuncular canal system.

<i>Aturia</i> Extinct genus of molluscs

Aturia is an extinct genus of Paleocene to Miocene nautilids within Aturiidae, a monotypic family, established by Campman in 1857 for Aturia Bronn, 1838, and is included in the superfamily Nautilaceae in Kümmel 1964.

Jovellania is a genus of extinct prehistoric nautiloids from the order Oncocerida known from the Lower Devonian of Europe. Nautiloids form a broad group of shelled cephalopods that were once diverse and numerous but are now represented by only a handful of species in two genera.

Brevicoceras is an extinct nautiloid genus from the order Oncocerida with wide distribution in the Middle Devonian in Eastern North America, Russia and Morocco. Nautiloids form a broad group of shelled cephalopods that were once diverse and numerous but are now represented by only a handful of species in two genera.

Westonoceras is an extinct nautiloid genus from the Discosorida that lived during the Middle and Late Ordovician that has been found in North America, Greenland, and Northern Europe. It is the type genus for the Westonoceratidae

<span class="mw-page-title-main">Orthoceratoidea</span> Extinct subclass of cephalopods

Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars, extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer.

Mandaloceratidae is a family in the nautiloid cephalopod order Discosorida, from the Middle and Upper(?) Silurian characterized by short, essentially straight shells referred to as breviconic, typically with a faintly exogastric shape produced by the profile of the body chamber.

<span class="mw-page-title-main">Endoceratidae</span> Extinct family of molluscs

Endoceratidae is a family of large to very large straight shelled nautiloid cephalopods belonging to the order Endocerida that lived during the Middle and Late Ordovician. They include the largest known Paleozoic invertebrates, represented by Endoceras and Cameroceras.

Pentameroceras is a straight to slightly exogastric breviconic oncocerid from the middle Silurian of North America and Europe belonging to the Trimeroceratidae.

Proterovaginoceras is a medium to large sized endocerid from the Early and Middle Ordovician included in the family Endoceratidae.

Simardoceras is a genus in the discosorid family Westonoceratidae from the Middle Ordovician of Quebec.

Discosoridae comprise a family of endogastric discosorids, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.

References

  1. "Fossil reveals giant underwater killing machine that once called outback Australia home". www.abc.net.au. 2021-07-11. Retrieved 2021-07-12.
  2. Teichert, C.; Kummel, B. (1960). "Size of endoceroid cephalopods". Breviora. 128: 1–7.
  3. Klug, C.; De Baets, K.; Kröger, B.; Bell, M.A.; Korn, D.; Payne, J.L. (2015). "Normal giants? Temporal and latitudinal shifts of Palaeozoic marine invertebrate gigantism and global change". Lethaia. 48 (2): 267–288. doi:10.1111/let.12104.
  4. Flower, R.H. (1955). "Status of endoceroid classification". Journal of Paleontology. 29 (3): 329–371. JSTOR   1300321.