Endoceratidae Temporal range: | |
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Fossil of Endoceras in Milan | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Cephalopoda |
Subclass: | Nautiloidea |
Order: | † Endocerida |
Family: | † Endoceratidae Hyatt, 1884 |
Endoceratidae is a family of large to very large straight shelled nautiloid cephalopods belonging to the order Endocerida that lived during the Middle and Late Ordovician. They include the largest known Paleozoic invertebrates, represented by Endoceras and Cameroceras . [1] [2]
Endocerids are characterized by large, generally ventral siphuncles with simple endocones which lack the more complex structures associated with the Proterocameroceratidae and their derivatives. [2] Endocones, nested cone-shaped deposits characteristic of the Endocerida, form in the apical portion of the siphuncle where they balanced the weight of the animal in the body chamber at the opposite end. This helped assure a horizontal orientation while the endoceratid animal was alive. The septal necks are holochoantitic, [2] reaching the previous septum and sometimes beyond, as macrochoantic. Connecting rings, which may be moderately thin, form a lining on the interior of the necks. [1] In some forms, such as Nanno and Chazyoceras the siphuncle is swollen at the apex so as to preclude the presence of adjacent camerae. In others, such as Cameroceras and Vaginoceras the siphuncle is tubular with chambers formed adjacently from the very start. The taxonomic significance of either in undetermined, other than as used to define a particular genus.
The Endoceratidae arose from the upper Canadian (Lower Ordovician) Piloceratidae according to Rousseau Flower, [3] [4] which have similar simple structured siphuncles, although they may resemble proterocameroceratids in general form. Accordingly, Flower [2] included the Endoceratdae and Piloceratidae, along with the Cyrtendoceratidae, in a suborder, the Endoceratina.
Flower restricted the Endoceratidae to those genera from the Middle and Upper Ordovician, and possibly Lower Silurian, belonging to the Endocerida with holochoantic or macrochoanitic septal necks and simple endocones. Teichert [1] added Proterovaginoceras , Triendoceras . and Allocotoceras from the Lower Ordovician
Genera included are:
Endocerida is an extinct nautiloid order, a group of cephalopods from the Lower Paleozoic with cone-like deposits in their siphuncle. Endocerida was a diverse group of cephalopods that lived from the Early Ordovician possibly to the Late Silurian. Their shells were variable in form. Some were straight (orthoconic), others curved (cyrtoconic); some were long (longiconic), others short (breviconic). Some long-shelled forms like Endoceras attained shell lengths close to 6 metres (20 ft). The related Cameroceras is anecdotally reported to have reached lengths approaching 9 metres (30 ft), but these claims are problematic. The overwhelming majority of endocerids and nautiloids in general are much smaller, usually less than a meter long when fully grown.
Cameroceras is a genus of extinct, large orthoconic cephalopod that lived mainly during the Ordovician period. It first appears during the middle Ordovician, around 470 million years ago, and was a fairly common component of the fauna in some places during the period, inhabiting the shallow seas of Laurentia, Baltica and Siberia. Its diversity and abundance became severely reduced following the Ordovician–Silurian extinction events, and the last remnants of the genus went extinct sometime during the Wenlock. C. rowenaense from Cincinnati had a shell length about 70 centimetres (2.3 ft), and unnamed species had length about 1 metre (3.3 ft). Although in older literature Cameroceras was treated as the largest nautiloid with shell length about 6 metres (20 ft) or even 30 feet (9.1 m), later study shows that the large-sized species no longer belongs to genus Cameroceras but Endoceras giganteum instead, and the 30 feet (9.1 m)-long specimen is highly doubtful.
Endoceras is an extinct genus of large, straight shelled cephalopods that gives its name to the Nautiloid order Endocerida. The genus lived during the middle and upper Ordovician 485 to 419 million years ago. The cross section in the mature portion is slightly wider than high, but is narrower laterally in the young. Sutures are straight and transverse. Endoceras has a large siphuncle, located close to the ventral margin, composed of concave segments, especially in the young but which may be tubular in the adult stage. Endocones are simple, subcircular in cross section, and penetrated by a narrow tube which may contain diaphragms reminiscent of the Ellesmerocerid ancestor.
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.
Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
The Proterocameroceratidae were the first of the Endocerida. They began early in the Ordovician with Proendoceras or similar genus which had developed endocones, replacing the diaphragms of the ellesmerocerid ancestor.
The Piloceratidae are a compressed, rapidly expanding, cyrtoconic brevicones with holochoanitic ventral siphuncles and simple endocones. Most likely evolved from Clitendoceras, a narrow, slightly endogastric genus intermediate in form between straight shelled Proendoceras and the bulkier Piloceratidae. Found in shallow carbonate marine sediments of Demingian through the Cassinian age,.
Clitendoceras is a genus of cephalopods in the order Endocerida from the Lower Ordovician with an elongate shell with a slight downward, endogastric, curvature and a siphuncle that lies along the ventral margin. Common for endocerids, the chambers are short and the septa close spaced with sutures sloping forward across the back of the shell. Septal necks are short in the young, lengthening in the adult. Endocones are simple, but with the ventral side of the last formed projecting forward. The endosiphotube running down the middle is arched on top and somewhat flat on the lower side.
Anthoceras is a genus of straight, annulated, proterocamerioceratids from the Lower Ordovician, found in North America, NW Australia, and Siberia. The cross section is circular, the siphuncle moderately large, and marginal. Segments are constricted ; septal necks hemichoantici to subholochoantic ; connecting rings thick. Endocones are long and slightly asymmetric.
The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic).
Winnipegoceras is an extinct nautiloid genus from the Ordovician belonging to the Order Discosorida.
Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.
The Armenoceratidae are a family of early Paleozoic nautiloid cephalopods belonging to the order Actinocerida.
Troedssonellidae is a family of orthoceroid cephalopods from the Ordovician, derived from rod-bearing Baltoceratidae, that have a continuous lining within the siphuncle that resembles very thin and slender endocones. Shells are generally slender and orthoconic. The siphuncle is central or subcentral, composed of straight or slightly expanded segments. Septal necks generally short and connecting rings are thin. Thin cameral deposits are known, which along with the position of the siphuncle and thin connecting rings distinguishes them from the endocerids in which they have been included.
Chazyoceras is a moderately large endocerid included in the Endoceratidae with a Nanno type apex and a ventral siphuncle with a holochoanitic wall, characteristic of the family. The siphuncle swelling at the apex is subtriangular in longitudinal profile. The endocones are of medium length.
Triendoceras is a genus of fairly large endocerid named by Flower (1958), included in the Endoceratidae by Teichert (1964), characterized by a holochoanitic ventral siphuncle in which the cross section through the endocones has an opening in the shape of an isosceles triangle with a sharp apex (corner) pointing down. Triendoceras is found in the upper Lower Ordovician of Quebec and New York in North America and possibly in Ohio, and in eastern Europe. The type species is T. montrealense.
Westonoceratidae are exogastric, mostly compressed, Discosorida of moderate size from the Middle Ordovician to the Lower Silurian.
Proterovaginoceras is a medium to large sized endocerid from the Early and Middle Ordovician included in the family Endoceratidae.
Discosoridae comprise a family of endogastric discosorids,, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.