Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day.
Endocerida is an extinct nautiloid order, a group of cephalopods from the Lower Paleozoic with cone-like deposits in their siphuncle. Endocerida was a diverse group of cephalopods that lived from the Early Ordovician possibly to the Late Silurian. Their shells were variable in form. Some were straight (orthoconic), others curved (cyrtoconic); some were long (longiconic), others short (breviconic). Some long-shelled forms like Endoceras attained shell lengths close to 6 metres (20 ft). The related Cameroceras is anecdotally reported to have reached lengths approaching 9 metres (30 ft), but these claims are problematic. The overwhelming majority of endocerids and nautiloids in general are much smaller, usually less than a meter long when fully grown.
Orthocerida, also known as the Michelinocerida, is an order of extinct orthoceratoid cephalopods that lived from the Early Ordovician possibly to the Late Triassic. A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous, and the Eocene fossil Antarcticeras is sometimes considered a descendant of the orthocerids although this is disputed. They were most common however from the Ordovician to the Devonian.
Endoceras is an extinct genus of large, straight shelled cephalopods that gives its name to the Nautiloid order Endocerida. The genus lived during the middle and upper Ordovician 485 to 419 million years ago. The cross section in the mature portion is slightly wider than high, but is narrower laterally in the young. Sutures are straight and transverse. Endoceras has a large siphuncle, located close to the ventral margin, composed of concave segments, especially in the young but which may be tubular in the adult stage. Endocones are simple, subcircular in cross section, and penetrated by a narrow tube which may contain diaphragms reminiscent of the Ellesmerocerid ancestor.
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.
The Piloceratidae are a compressed, rapidly expanding, cyrtoconic brevicones with holochoanitic ventral siphuncles and simple endocones. Most likely evolved from Clitendoceras, a narrow, slightly endogastric genus intermediate in form between straight shelled Proendoceras and the bulkier Piloceratidae. Found in shallow carbonate marine sediments of Demingian through the Cassinian age,.
Meniscoceras is a straight and slender Chazyan endocerid described by Rousseau Flower in 1941 The genus was originally included in the Proterocameroceratidae but later placed with its predecessor, Najaceras, in the Najaceratidae.
Clitendoceras is a genus of cephalopods in the order Endocerida from the Lower Ordovician with an elongate shell with a slight downward, endogastric, curvature and a siphuncle that lies along the ventral margin. Common for endocerids, the chambers are short and the septa close spaced with sutures sloping forward across the back of the shell. Septal necks are short in the young, lengthening in the adult. Endocones are simple, but with the ventral side of the last formed projecting forward. The endosiphotube running down the middle is arched on top and somewhat flat on the lower side.
Mcqueenoceras is an extinct genus of early endocerid, a nautiloid from the Floian epoch of the late early Ordovician period. It was similar in overall form to Clitendoceras, from which it may have been derived. Mcqueenoceras, like Clitendoceras, has ventral siphuncle but the endocones are thicker on the ventral side and thinner on the dorsal. Also the sutures in Mcqueenoceras retreat rearward, forming lobes as they cross the venter. The type species is Mcqueenoceras jeffersonense, named by E.O. Ulrich and A.F. Foerste in 1935, and it is known from Missouri and New York. In 1956, Rousseau H. Flower named two other species, M. cariniferum and M. ventrale, both known from Maryland.
Allotrioceras is a tubular fossil from the Middle Ordovician of the state of New York, collected by Rousseau H. Flower; included by him in the Endocerida and placed in a new family, the Allotrioceratidae. Allotrioceras is characterized by a lateral pair of subequal conical structures, resembling the endocones of endocerids, separated by a straight partition that extends more than half way across from either the dorsal or ventral side, as perceived, and runs along the length. What remains is thought to represent a siphuncle, which ranges from about 8 millimeters (0.3 in) to about 15 millimeters (0.6 in) in diameter. On the basis of diagnosis, inclusion of Allotrioceras and the Allotrioceratidae, for which it is the type, in the endocerida seems at best tentative.
The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.
Troedssonellidae is a family of orthoceroid cephalopods from the Ordovician, derived from rod-bearing Baltoceratidae, that have a continuous lining within the siphuncle that resembles very thin and slender endocones. Shells are generally slender and orthoconic. The siphuncle is central or subcentral, composed of straight or slightly expanded segments. Septal necks generally short and connecting rings are thin. Thin cameral deposits are known, which along with the position of the siphuncle and thin connecting rings distinguishes them from the endocerids in which they have been included.
Campendoceras is a genus of proterocameroceratids from the Lower Ordovician of NW Australia and possibly Estonia that can be recognized by its slender, weakly endogastric, longiconic and annulate shell and large marginal siphuncle that contains endocones.
Cotteroceras is a genus of proterocameroceratids from the Lower Ordovician of North America and Siberia characterized by a long straight and compressed shell with very short camerae and long body chamber. Sutures are straight and oblique, sloping dorsoventrally toward the apex. The siphuncle is large but details are unknown.
Endoceratidae is a family of large to very large straight shelled nautiloid cephalopods belonging to the order Endocerida that lived during the Middle and Late Ordovician. They include the largest known Paleozoic invertebrates, represented by Endoceras and Cameroceras.
Chazyoceras is a moderately large endocerid included in the Endoceratidae with a Nanno type apex and a ventral siphuncle with a holochoanitic wall, characteristic of the family. The siphuncle swelling at the apex is subtriangular in longitudinal profile. The endocones are of medium length.
Triendoceras is a genus of fairly large endocerid named by Flower (1958), included in the Endoceratidae by Teichert (1964), characterized by a holochoanitic ventral siphuncle in which the cross section through the endocones has an opening in the shape of an isosceles triangle with a sharp apex (corner) pointing down. Triendoceras is found in the upper Lower Ordovician of Quebec and New York in North America and possibly in Ohio, and in eastern Europe. The type species is T. montrealense.
Perkinsocerasis an endocerid genus from the Middle Ordovician (Chazyan) of Champlain Valley established by Flower in 1976, which he added to his Allotrioceratidae based on certain similarities to Williamsoceras and Cacheoceras which had been added previously.
Proterocameroceras is an early Endocerid from the upper Lower Ordovician belong to the Proterocameroceratidae, characterized by a rather large, straight, longiconic shell, short chambers, generally straight sutures, and large ventral siphuncle with short septal necks, thick complex connecting rings, and endocones with three endosiphuncular blades toward the apex.
Westonoceratidae are exogastric, mostly compressed, Discosorida of moderate size from the Middle Ordovician to the Lower Silurian.
