Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting tail-like abdomen, usually hidden entirely under the thorax. They live in all the world's oceans, in freshwater, and on land. They are generally covered with a thick exoskeleton. They generally have five pairs of legs, and they have pincer claws on the ends of the frontmost pair. They first appeared during the Jurassic period, around 200 million years ago.
Hermit crabs are anomuran decapod crustaceans of the superfamily Paguroidea that have adapted to occupy empty scavenged mollusc shells to protect their fragile exoskeletons. There are over 800 species of hermit crab, most of which possess an asymmetric abdomen concealed by a snug-fitting shell. Hermit crabs' soft (non-calcified) abdominal exoskeleton means they must occupy shelter produced by other organisms or risk being defenseless.
The Japanese spider crab is a species of marine crab and is the biggest one that lives in the waters around Japan. At around 3.7 meters, it has the largest leg-span of any arthropod. The Japanese name for this species is taka-ashi-gani,, literally translating to “tall legs crab”. It goes through three main larval stages along with a prezoeal stage to grow to its great size.
The Dungeness crab makes up one of the most important seafood industries along the west coast of North America. Its typical range extends from Alaska's Aleutian Islands to Point Conception, near Santa Barbara, California. Dungeness typically grow 6–7 in (150–180 mm) at their widest point and inhabit eelgrass beds and sandy bottoms. Its common name comes from the Dungeness Spit in the Strait of Juan de Fuca, Washington state, United States, which shelters a shallow bay inhabited by the crabs.
Carcinus maenas is a common littoral crab. It is known by different names around the world. In the British Isles, it is generally referred to as the shore crab, or green shore crab. In North America and South Africa, it bears the name European green crab.
Portunus pelagicus, also known as the blue crab, blue swimmer crab, blue manna crab and flower crab is a species of large crab found in the Indo-Pacific, including off the coasts Indonesia, Malaysia, Cambodia, Thailand, the Philippines, and Vietnam; and in the intertidal estuaries around most of Australia and east to New Caledonia.
Mictyris is a genus of brightly coloured crabs, placed in its own taxonomical family, the Mictyridae. It inhabits the central Indo-West Pacific region. These crabs congregate on mud flats or beaches in groups of a few thousand, and filter sand or mud for microscopic organisms. They congregate during low tide, and bury themselves in the sand during high tide or whenever they are threatened. This is done in wet sand, and they dig in a corkscrew pattern, leaving many small round pellets of sand behind them.
Xanthoidea is a superfamily of crabs, comprising seven families. Formerly, a number of other families were included in Xanthoidea, but many of these have since been removed to other superfamilies. These include Carpilioidea, Eriphioidea, Hexapodoidea, Pilumnoidea and Trapezioidea. Even in this reduced state, Xanthoidea remains one of the most species-rich superfamilies of crabs.
The Gecarcinidae, the land crabs, are a family of true crabs that are adapted for terrestrial existence. Similar to all other crabs, land crabs possess a series of gills. In addition, the part of the carapace covering the gills is inflated and equipped with blood vessels. These organs extract oxygen from the air, analogous to the vertebrate lungs. Adult land crabs are terrestrial, but visit the sea periodically, where they breed and their larvae develop. Land crabs are tropical omnivores which sometimes cause considerable damage to crops. Most land crabs have one of their claws larger than the other.
Cardisoma is a genus of large land crabs. Three species formerly placed in this genus are now placed in Discoplax. The four species that remain in Cardisoma are found in warm coastal regions where they live in burrows. Young individuals are often very colourful with a purple-blue carapace and orange-red legs, but as they grow older the colours tend to fade, and females may be duller than males. Although less extreme than in fiddler crabs, one claw is usually considerably larger than the other. They are omnivores, but primarily feed on plant material.
Gecarcinucoidea is a superfamily of freshwater crabs. Its members have been grouped into families in various ways, with some authors recognizing families such as "Deckeniidae", "Sundathelphusidae", and "Parathelphusidae", but now only the family Gecarcinucidae is currently recognized.
Chiromantes haematocheir, commonly known as red-clawed crab, is a species of mudflat crab in the family Sesarmidae. It is endemic to East Asia and is quite distinct from the other species placed in the genus Chiromantes. The genus may be restricted to this one species.
The coconut crab is a terrestrial species of giant hermit crab, and is also known as the robber crab or palm thief. It is the largest terrestrial arthropod known, with a weight of up to 4.1 kg (9 lb). The distance from the tip of one leg to the tip of another can be as wide as 1 m. It is found on islands across the Indian and Pacific Oceans, as far east as the Gambier Islands, Pitcairn Islands and Caroline Island and as far west as Zanzibar. While its range broadly shadows the distribution of the coconut palm, the coconut crab has been extirpated from most areas with a significant human population such as mainland Australia and Madagascar.
Eubrachyura is a group of decapod crustaceans comprising the more derived crabs. It is divided into two subsections, based on the position of the genital openings in the two sexes. In the Heterotremata, the openings are on the legs in the males, but on the sternum in females, while in the Thoracotremata, the openings are on the sternum in both sexes. This contrasts with the situation in other decapods, in which the genital openings are always on the legs. Heterotremata is the larger of the two groups, containing the species-rich superfamilies Xanthoidea and Pilumnoidea and all the freshwater crabs. The eubrachyura is well known for actively and constantly building its own burrows. The fossil record of the Eubrachyura extends back to the Cretaceous; the supposed Bathonian representative of the group, Hebertides jurassica, ultimately turned out to be Cenozoic in age.
Heterotremata is a clade of crabs, comprising those crabs in which the genital openings are on the sternum in females, but on the legs in males. It comprises 68 families in 28 superfamilies.
Cheiragonidae is a small family of crabs, sometimes called helmet crabs, placed in its own superfamily, Cheriagonoidea. It comprises three extant species, Erimacrus isenbeckii, Telmessus acutidens and Telmessus cheiragonus, there are no yet evidences of Cheiragonidae in the fossil record. Many of these crabs were formerly treated as members of the Atelecyclidae.
Matutidae is a family of crabs, sometimes called moon crabs, adapted for swimming or digging. They differ from the swimming crabs of the family Portunidae in that all five pairs of legs are flattened, rather than just the last pair, as in Portunidae. Crabs in the Matutidae are aggressive predators.
Gecarcinus ruricola is a species of terrestrial crab. It is the most terrestrial of the Caribbean land crabs, and is found from western Cuba across the Antilles as far east as Barbados. Common names for G. ruricola include the purple land crab, black land crab, red land crab, and zombie crab.
Aratus pisonii, commonly known as the mangrove tree crab, is a species of crab which lives in mangrove trees in tropical and subtropical parts of the Americas, from Florida to Brazil on the Atlantic coast. A.pacificus occurs from Nicaragua to Peru on the Pacific coast. A. pisoni feeds mostly on the leaves of the mangroves, but is an omnivore, and prefers animal matter when possible. A. pisonii and A. pacificus are the two species in the genus Aratus. The specific epithet pisonii commemorates the Dutch naturalist Willem Piso who travelled in Brazil in 1638 with Georg Marggraf.
Epialtidae is a family of crabs, containing the subfamilies:
