Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen), usually hidden entirely under the thorax. They live in all the world's oceans, in freshwater, and on land, are generally covered with a thick exoskeleton, and have a single pair of pincers on each arm. They first appeared during the Jurassic Period.
Hermit crabs are anomuran decapod crustaceans of the superfamily Paguroidea that have adapted to occupy empty scavenged mollusc shells to protect their fragile exoskeletons. There are over 800 species of hermit crab, most of which possess an asymmetric abdomen concealed by a snug-fitting shell. Hermit crabs' soft (non-calcified) abdominal exoskeleton means they must occupy shelter produced by other organisms or risk being defenseless.
The Decapoda or decapods are an order of crustaceans within the class Malacostraca, and includes crabs, lobsters, crayfish, shrimp, and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 extant species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp and Anomura including hermit crabs, porcelain crabs, squat lobsters making up the bulk of the remainder. The earliest fossils of the group date to the Devonian.
The fiddler crab or calling crab may be any of more than one hundred species of semiterrestrial marine crabs in the family Ocypodidae, well known for their sexually dimorphic claws; the males' major claw is much larger than the minor claw, while the females' claws are both the same size. A smaller number of ghost crab and mangrove crab species are also found in the family Ocypodidae. This entire group is composed of small crabs, the largest being slightly over two inches (5 cm) across. Fiddler crabs are found along sea beaches and brackish intertidal mud flats, lagoons, swamps, and various other types of brackish or salt-water wetlands.
The Japanese spider crab is a species of marine crab and is the biggest one that lives in the waters around Japan. It has the largest known leg-span of any arthropod. The Japanese name for this species is taka-ashi-gani,, literally translating to “tall legs crab”. It goes through three main larval stages along with a prezoeal stage to grow to its great size.
The Dungeness crab is a species of crab inhabiting eelgrass beds and water bottoms along the west coast of North America. It typically grows to 20 cm (7.9 in) across the carapace and is a popular seafood. Its common name comes from Dungeness Spit, United States, which shelters a shallow bay inhabited by the crabs.
Carcinus maenas is a common littoral crab. It is known by different names around the world. In the British Isles, it is generally referred to as the shore crab, or green shore crab. In North America and South Africa, it bears the name European green crab.
Portunus pelagicus, also known as the blue crab, blue swimmer crab, blue manna crab and flower crab is a species of large crab found in the Indo-Pacific, including off the coasts Indonesia, Malaysia, Cambodia, Thailand, the Philippines, and Vietnam; and in the intertidal estuaries around most of Australia and east to New Caledonia.
Mictyris is a genus of brightly coloured crabs, placed in its own taxonomical family, the Mictyridae. It inhabits the central Indo-West Pacific region. These crabs congregate on mud flats or beaches in groups of a few thousand, and filter sand or mud for microscopic organisms. They congregate during low tide, and bury themselves in the sand during high tide or whenever they are threatened. This is done in wet sand, and they dig in a corkscrew pattern, leaving many small round pellets of sand behind them.
Xanthoidea is a superfamily of crabs, comprising the three families Xanthidae, Panopeidae and Pseudorhombilidae. Formerly, a number of other families were included in Xanthoidea, but many of these have since been removed to other superfamilies. These include Carpilioidea, Eriphioidea, Hexapodoidea, Pilumnoidea and Trapezioidea. Even in this reduced state, Xanthoidea remains one of the most species-rich superfamilies of crabs.
The Gecarcinidae, the land crabs, are a family of true crabs that are adapted for terrestrial existence. Similar to all other crabs, land crabs possess a series of gills. In addition, the part of the carapace covering the gills is inflated and equipped with blood vessels. These organs extract oxygen from the air, analogous to the vertebrate lungs. Adult land crabs are terrestrial, but visit the sea periodically, where they breed and their larvae develop. Land crabs are tropical omnivores which sometimes cause considerable damage to crops. Most land crabs have one of their claws larger than the other.
Cardisoma is a genus of large land crabs. Three species formerly placed in this genus are now placed in Discoplax. The four species that remain in Cardisoma are found in warm coastal regions where they live in burrows. Young individuals are often very colourful with a purple-blue carapace and orange-red legs, but as they grow older the colours tend to fade, and females may be duller than males. Although less extreme than in fiddler crabs, one claw is usually considerably larger than the other. They are omnivores, but primarily feed on plant material.
Gecarcinucoidea is a superfamily of freshwater crabs. Its members have been grouped into families in various ways, with some authors recognizing families such as "Deckeniidae", "Sundathelphusidae", and "Parathelphusidae", but now only the family Gecarcinucidae is currently recognized.
Lybia is a genus of small crabs in the family Xanthidae. Their common names include boxer crabs, boxing crabs and pom-pom crabs. They are notable for their mutualism with sea anemones, which they hold in their claws for defense. In return, the anemones get carried around, which may enable them to capture more food particles with their tentacles. Boxer crabs use at least three species of anemones, including Bundeopsis spp. and Triactis producta. The bonding with the anemone is not needed for survival, however, and boxer crabs have frequently been known to live without them, sometimes substituting other organisms such as sponges and corals for the sea anemones.
Eubrachyura is a group of decapod crustaceans comprising the more derived crabs. It is divided into two subsections, based on the position of the genital openings in the two sexes. In the Heterotremata, the openings are on the legs in the males, but on the sternum in females, while in the Thoracotremata, the openings are on the sternum in both sexes. This contrasts with the situation in other decapods, in which the genital openings are always on the legs. Heterotremata is the larger of the two groups, containing the species-rich superfamilies Xanthoidea and Pilumnoidea and all the freshwater crabs. The eubrachyura is well known for actively and constantly building its own burrows. The fossil record of the Eubrachyura extends back to the Cretaceous; the supposed Bathonian representative of the group, Hebertides jurassica, ultimately turned out to be Cenozoic in age.
Heterotremata is a clade of crabs, comprising those crabs in which the genital openings are on the sternum in females, but on the legs in males. It comprises 68 families in 28 superfamilies.
Cheiragonidae is a small family of crabs, sometimes called helmet crabs, placed in its own superfamily, Cheriagonoidea. It comprises three extant species, Erimacrus isenbeckii, Telmessus acutidens and Telmessus cheiragonus, there are no yet evidences of Cheiragonidae in the fossil record. Many of these crabs were formerly treated as members of the Atelecyclidae.
Gecarcinus ruricola is a species of terrestrial crab. It is the most terrestrial of the Caribbean land crabs, and is found from western Cuba across the Antilles as far east as Barbados. Common names for G. ruricola include the purple land crab, black land crab, red land crab, and zombie crab.
Aratus pisonii, commonly known as the mangrove tree crab, is a species of crab which lives in mangrove trees in tropical and subtropical parts of the Americas, from Florida to Brazil on the Atlantic coast. A.pacificus occurs from Nicaragua to Peru on the Pacific coast. A. pisoni feeds mostly on the leaves of the mangroves, but is an omnivore, and prefers animal matter when possible. A. pisonii and A. pacificus are the two species in the genus Aratus. The specific epithet pisonii commemorates the Dutch naturalist Willem Piso who travelled in Brazil in 1638 with Georg Marggraf.
Epialtidae is a family of crabs, containing the subfamilies:
