Portunidae is a family of crabs which contains the swimming crabs.
Cancridae is a family of crabs. It comprises six extant genera, and eleven exclusively fossil genera, in two subfamilies:
Xanthoidea is a superfamily of crabs, comprising the three families Xanthidae, Panopeidae and Pseudorhombilidae. Formerly, a number of other families were included in Xanthoidea, but many of these have since been removed to other superfamilies. These include Carpilioidea, Eriphioidea, Hexapodoidea, Pilumnoidea and Trapezioidea. Even in this reduced state, Xanthoidea remains one of the most species-rich superfamilies of crabs.
Retroplumidae is a family of heterotrematan crabs, placed in their own (monotypic) superfamily, Retroplumoidea.
The thumbnail crab, Thia scutellata, is a species of crab whose carapace resembles a human thumbnail. It is found in the North Sea, north-east Atlantic and Mediterranean Sea. It is the only extant species in the genus Thia, although two fossil species are known.
The Varunidae are a family of thoracotrematan crabs. The delimitation of this family, part of the taxonomically confusing Grapsoidea, is undergoing revision. For a long time, they were placed at the rank of subfamily in the Grapsidae, but they appear to be closest to Macropthalmus and the Mictyridae, which are usually placed in the Ocypodoidea. It may thus be better to merge the latter superfamily with the Grapsoidea, retaining the latter name as it is older.
Goneplacidae is a family of crabs of the order Decapoda and the superfamily Goneplacoidea. It includes the following genera:
Hexapodidae is a family of crabs, the only family in the superfamily Hexapodoidea. It has traditionally been treated as a subfamily of the family Goneplacidae, and was originally described as a subfamily of Pinnotheridae. Its members can be distinguished from all other true crabs by the reduction of the thorax, such that only seven sternites are exposed, and only four pairs of pereiopods are present. Not counting the enlarged pair of claws, this leaves only six walking legs, from which the type genus Hexapus, and therefore the whole family, takes its name. Some anomuran "crabs", such as porcelain crabs and king crabs also have only four visible pairs of legs. With the exception of Stevea williamsi, from Mexico, all the extant members are found either in the Indo-Pacific oceans, or around the coast of Africa.
Brachyura is a group of crustaceans comprising the more derived crabs. It is divided into two subsections, based on the position of the genital openings in the two sexes. In the Heterotremata, the openings are on the legs in the males, but on the sternum in females, while in the Thoracotremata, the openings are on the sternum in both sexes. This contrasts with the situation in other decapods, in which the genital openings are always on the legs. Heterotremata is the larger of the two groups, containing the species-rich superfamilies Xanthoidea and Pilumnoidea and all the freshwater crabs (Gecarcinucoidea, Potamoidea. The eubrachyura is well known for actively and constantly building its own burrows. The fossil record of the Eubrachyura extends back to the Cretaceous; the supposed Bathonian representative of the group, Hebertides jurassica, ultimately turned out to be Cenozoic in age.
Cheiragonidae is a small family of crabs, sometimes called helmet crabs, placed in its own superfamily, Cheriagonoidea. It comprises three extant species, Erimacrus isenbeckii, Telmessus acutidens and Telmessus cheiragonus, there are no yet evidences of Cheiragonidae in the fossil record. Many of these crabs were formerly treated as members of the Atelecyclidae.
Pilumnoidea is a superfamily of crabs, whose members were previously included in the Xanthoidea. The three families are unified by the free articulation of all the segments of the male crab's abdomen and by the form of the gonopods. The earliest fossils assigned to this group are of Eocene age.
Actumnus is a genus of crabs in the family Pilumnidae. Alongside the 28 extant species, it has a fossil record extending back into the Miocene.
Palicoidea is a superfamily of crabs, comprising the two families Crossotonotidae and Palicidae. Together, they contain 13 genera, including two genera in the Palicidae known only from fossils. The two families were previously treated as two subfamilies in a Palicidae of wider circumscription.
Epialtidae is a family of crabs, containing the subfamilies:
Xantho is a genus of crabs in the family Xanthidae, containing five extant species, all restricted to the north-east Atlantic Ocean and Mediterranean Sea, although Xantho granulicarpis is not universally recognised as a separate species from Xantho hydrophilus:
Xanthodius is a genus of crabs in the family Xanthidae, containing one exclusively fossil species and the following extant species:
Zosimus is a genus of crabs in the family Xanthidae, containing the following species:
Mathildellidae is family of crabs belonging to the superfamily Goneplacoidea, containing the following genera:
Dynomenidae is a family of crabs in the superfamily Dromioidea mostly found in Madagascar. There are nineteen genera in this family: five extant and fourteen known from fossils:
Ethusidae is one of two extant families of crabs in the superfamily Dorippoidea. It contains four genera. Members of this family are found in marine environments at depths from 16m to 4,192m.
