Crinoids are marine invertebrates that make up the class Crinoidea. Crinoids that remain attached to the sea floor by a stalk in their adult form are commonly called sea lilies, while the unstalked forms, called feather stars or comatulids, are members of the largest crinoid order, Comatulida. Crinoids are echinoderms in the phylum Echinodermata, which also includes the starfish, brittle stars, sea urchins and sea cucumbers. They live in both shallow water and in depths as great as 9,000 meters (30,000 ft).
Arkarua adami is a small, Precambrian disk-like fossil with a raised center, a number of radial ridges on the rim, and a five-pointed central depression marked with radial lines of five small dots from the middle of the disk center. Fossils range from 3 to 10 mm in diameter.
Blastoids are an extinct type of stemmed echinoderm, often referred to as sea buds. They first appear, along with many other echinoderm classes, in the Ordovician period, and reached their greatest diversity in the Mississippian subperiod of the Carboniferous period. However, blastoids may have originated in the Cambrian. Blastoids persisted until their extinction at the end of Permian, about 250 million years ago. Although never as diverse as their contemporary relatives, the crinoids, blastoids are common fossils, especially in many Mississippian-age rocks.
Edrioasteroidea is an extinct class of echinoderms. The living animal would have resembled a pentamerously symmetrical disc or cushion. They were obligate encrusters and attached themselves to inorganic or biologic hard substrates. A 507 million years old species, Totiglobus spencensis, is actually the first known echinoderm adapted to live on a hard surface after the soft microbial mats that covered the seafloor were destroyed in the Cambrian substrate revolution.
Helicoplacoidea is an extinct class within the Echinodermata. All known taxa were discovered in sediments dating back to the Cambrian.
Marine invertebrates are the invertebrates that live in marine habitats. Invertebrate is a blanket term that includes all animals apart from the vertebrate members of the chordate phylum. Invertebrates lack a vertebral column, and some have evolved a shell or a hard exoskeleton. As on land and in the air, marine invertebrates have a large variety of body plans, and have been categorised into over 30 phyla. They make up most of the macroscopic life in the oceans.
The "Cambrian substrate revolution" or "Agronomic revolution", evidenced in trace fossils, is a sudden diversification of animal burrowing during the early Cambrian period.
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the late Ediacaran and early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span.
The Cambrian explosion is an interval of time approximately 538.8 million years ago in the Cambrian period of the early Paleozoic when a sudden radiation of complex life occurred, and practically all major animal phyla started appearing in the fossil record. It lasted for about 13 to 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.
Deuterostomes are bilaterian animals of the superphylum Deuterostomia, typically characterized by their anus forming before the mouth during embryonic development. Deuterostomia is further divided into 4 phyla: Chordata, Echinodermata, Hemichordata, and the extinct Vetulicolia known from Cambrian fossils. The extinct clade Cambroernida is also thought to be a member of Deuterostomia.
Camptostroma roddyi is an extinct echinoderm from the Bonnia-Olenellus Zone the Early Cambrian Kinzers Formation near York and Lancaster, Southeastern Pennsylvania. In life, it would have resembled a cupcake, with the axial skeleton forming a star pattern on the upper surface. It was originally thought, on the basis of its medusoid shape, to be a jellyfish-like organism, but the fossils themselves clearly rule out the possibility of a gelatinous body - the stereom plates are clearly preserved and possess the calcitic cleavage pattern diagnostic of echinoderms. It has been placed in a class of basal echinoderms, the Edrioasteroids.
Homalozoa is an obsolete extinct subphylum of Paleozoic era echinoderms, prehistoric marine invertebrates. They are also referred to as carpoids.
Walcottidiscus is a genus of pentagonally shaped echinoderms known from the Middle Cambrian Burgess Shale. 16 specimens of Walcottidiscus are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community.
The Fezouata Formation or Fezouata Shale is a geological formation in Morocco which dates to the Early Ordovician. It was deposited in a marine environment, and is known for its exceptionally preserved fossils, filling an important preservational window beyond the earlier and more common Cambrian Burgess shale-type deposits. The fauna of this geological unit is often described as the Fezouata biota, and the particular strata within the formation which exhibit exceptional preservation are generally termed the Fezouata Lagerstätte.
Stereom is a calcium carbonate material that makes up the internal skeletons found in all echinoderms, both living and fossilized forms. It is a sponge-like porous structure which, in a sea urchin may be 50% by volume living cells, and the rest being a matrix of calcite crystals. The size of openings in stereom varies in different species and in different places within the same organism. When an echinoderm becomes a fossil, microscopic examination is used to reveal the structure and such examination is often an important tool to classify the fossil as an echinoderm or related creature.
Cincta is an extinct class of echinoderms that lived only in the Middle Cambrian epoch. Homostelea is a junior synonym. The classification of cinctans is controversial, but they are probably part of the echinoderm stem group.
Soluta is an extinct class of echinoderms that lived from the Middle Cambrian to the Early Devonian. The class is also known by its junior synonym Homoiostelea. Soluta is one of the four "carpoid" classes, alongside Ctenocystoidea, Cincta, and Stylophora, which made up the obsolete subphylum Homalozoa. Solutes were asymmetric animals with a stereom skeleton and two appendages, an arm extending anteriorly and a posterior appendage called a homoiostele.
Ctenocystoidea is an extinct clade of echinoderms, which lived during the Cambrian and Ordovician periods. Unlike other echinoderms, ctenocystoids had bilateral symmetry, or were only very slightly asymmetrical. They are believed to be one of the earliest-diverging branches of echinoderms, with their bilateral symmetry a trait shared with other deuterostomes. Ctenocystoids were once classified in the taxon Homalozoa, also known as Carpoidea, alongside cinctans, solutes, and stylophorans. Homalozoa is now recognized as a polyphyletic group of echinoderms without radial symmetry. Ctenocystoids were geographically widespread during the Middle Cambrian, with one species surviving into the Late Ordovician.
Yanjiahella biscarpa is an extinct species of Early Cambrian deuterostome which may represent the earliest stem group echinoderm.
Yorkicystis is a genus of edrioasteroid echinoderm that lived 510 million years ago in the Cambrian aged Kinzers Formation in what is now Pennsylvania. This genus is important as it provides some of the oldest evidence of echinoderms losing their hard mineralized outer skeletons. Yorkicystis also shows that some echinoderms lost their skeletons during the Cambrian, which is a greatly different time as to when most other species lost theirs.
