Helsby Sandstone Formation

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Helsby Sandstone Formation
Stratigraphic range: Middle Triassic Anisian
Ladram Bay , Ladram Rock ^ Point - geograph.org.uk - 1831352.jpg
Exposures of Otter Sandstone at Ladram Bay, Devon, England
Type Formation
Unit of Sherwood Sandstone Group (SSG)
Overlies Wilmslow Sandstone Formation (disconformity) or Chester Formation (unconformity)
ThicknessUp to 500 m
Lithology
Primary sandstone
Other siltstone, mudstone, conglomerate
Location
Region England
Country United Kingdom

The Helsby Sandstone Formation, formerly known as the Otter Sandstone (in Devon and Somerset), is a geological formation in England. It was deposited during the Anisian stage of the Middle Triassic epoch. It primarily consists of medium to fine grained sandstone, both cross bedded and flat bedded, with minor lenticular beds of siltstone and mudstone and thin beds of conglomerate in some areas. It is found across England, from Devon on the southern coast to Cumbria in the north, with some exposures in Worcestershire and in Staffordshire in the Midlands. [1] It was deposited in a hot, arid climate, when Britain formed part of the Pangean interior [2] [3] with the cross-bedded sandstones of the formation representing remnants of wind-blown sand dunes, [4] while the majority of the formation was produced by the action of braided rivers, specifically the ‘Budleighensis’ river system, which flowed northwards from what is now northern France through southern and central England, before draining into what is now the Irish Sea. The environment of the formation likely also included ponds and small lakes, with surrounding vegetation including conifers (whose roots are preserved as concretions) and horsetails. Towards the top of the formation, the conditions become more arid, with the overlying formations showing the disappearance of rivers and the existence of playa-like conditions. [3]

The formation in its southern coastal exposure in Devon in the vicinity of Sidmouth has provided important fossils of contemporary vertebrates, including temnospondyl amphibians, procolophonid, rhynchosaurian, sphenodontian and archosaurian reptiles, as well as fish. [3]

Paleobiota

Taken from [3] unless otherwise noted:

Fish of the Helsby Sandstone
GenusSpeciesNotesImage
Dipteronotus D. cyphusA small 10–12 centimetres (3.9–4.7 in) long perleidiform fish, known from complete specimens, most common fish in the formation. [5]
Fossil of the closely related species D. aculeatus Dipteronotus aculeatus.jpg
Fossil of the closely related species D. aculeatus
Actinopterygii IndeterminateIncludes a fish known from a single mostly complete 10–12 centimetres (3.9–4.7 in) long undescribed specimen distinct from Dipteronotus, as well as the jaw of a pike-like fish, as well as other indeterminate microvertebrate remains. [5]
Elasmobranchii IndeterminateFreshwater shark known from a fin spine
Amphibians of the Helsby Sandstone
GenusSpeciesNotesImage
Eocyclotosaurus A capitosaurian temnospondyl
Eocyclotosaurus wellesi1DB.jpg
Mastodonsaurus?A capitosaurian temnospondyl, some authors have consider the remains indeterminate and not belonging to the genus [3]
Mastodontosaurus spp DB24.jpg
Temnospondyli IndeterminateIncludes a third taxon apparently distinct from Eocyclotosaurus and Mastodonsaurus, as well as indeterminate fragmentary material
Reptiles of the Helsby Sandstone
GenusSpeciesNotesImage
Kapes K. bentoniA small lizard-like procolophonid reptile, known from a partial skeleton
Kapes full skeleton.jpg
Agriodontosaurus [6] A. helsbypetraeA sphenodontian reptile known from a partial skeleton
Agriodontosaurus (holotype, BRSUG 29950-14).png
Fodonyx F. spenceriA rhynchosaurian reptile, known from postcranial and skull material
Fodonyx NT.jpg
Bentonyx B. sidensisA rhynchosaurian reptile, known from a skull
Bentonyx sidensis.jpg
Feralisaurus [7] F. coramiA small lepidosauromorph known from specimen BRSUG 29950-12, an articulated skeleton
Coartaredens C. isaaciA reptile of uncertain classifcation, possibly a lepidosauromorph, but this is uncertain as it is only known from fragmentary remains
Chirotherium Ichnogenus of fossil trackways thought to be produced by "rauisuchians"
Archosauria IndeterminateRemains include vertebrae, teeth and a jaw fragment, which may represent poposauroids, with the jaw fragment suggested to possibly represent Bromsgroveia.
Rhynchosauria Most common vertebrates in the formation, remains include a headless skeleton
Procolophonidae Several indeterminate procolophonids distinct from Kapes, one of which has similarities to Sclerosaurus

References

  1. "Helsby Sandstone Formation- BGS Lexicon of Named Rock Units". webapps.bgs.ac.uk. Retrieved 2025-09-10.
  2. Bloomfield, J. P.; Moreau, M. F.; Newell, A. J. (January 2006). "Characterization of permeability distributions in six lithofacies from the Helsby and Wilmslow sandstone formations of the Cheshire Basin, UK". Geological Society, London, Special Publications. 263 (1): 83–101. Bibcode:2006GSLSP.263...83B. doi:10.1144/GSL.SP.2006.263.01.04. ISSN   0305-8719.
  3. 1 2 3 4 5 Coram, Robert A.; Radley, Jonathan D.; Benton, Michael J. (June 2019). "The Middle Triassic (Anisian) Otter Sandstone biota (Devon, UK): review, recent discoveries and ways ahead" . Proceedings of the Geologists' Association. 130 (3–4): 294–306. Bibcode:2019PrGA..130..294C. doi:10.1016/j.pgeola.2017.06.007.
  4. Mountney, Nigel P.; Thompson, David B. (August 2002). "Stratigraphic evolution and preservation of aeolian dune and damp/wet interdune strata: an example from the Triassic Helsby Sandstone Formation, Cheshire Basin, UK" . Sedimentology. 49 (4): 805–833. Bibcode:2002Sedim..49..805M. doi:10.1046/j.1365-3091.2002.00472.x. ISSN   0037-0746.
  5. 1 2 Coram, Robert A.; Radley, Jonathan D.; Benton, Michael J. (September 2021). "Triassic tragedy—a bone bed in the Otter Sandstone of East Devon, south-west England" . Geology Today. 37 (5): 176–183. Bibcode:2021GeolT..37..176C. doi:10.1111/gto.12367. ISSN   0266-6979.
  6. Marke, Daniel; Whiteside, David I.; Sethapanichsakul, Thitiwoot; Coram, Robert A.; Fernandez, Vincent; Liptak, Alexander; Newham, Elis; Benton, Michael J. (2025-09-10). "The oldest known lepidosaur and origins of lepidosaur feeding adaptations". Nature: 1–10. doi: 10.1038/s41586-025-09496-9 . ISSN   0028-0836.
  7. Cavicchini, Iacopo; Zaher, Marta; Benton, Michael J. (2020-05-03). "An Enigmatic Neodiapsid Reptile from the Middle Triassic of England". Journal of Vertebrate Paleontology. 40 (3) e1781143. Bibcode:2020JVPal..40E1143C. doi: 10.1080/02724634.2020.1781143 . ISSN   0272-4634.