Hippopotamus gorgops Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Family: | Hippopotamidae |
Genus: | Hippopotamus |
Species: | †H. gorgops |
Binomial name | |
†Hippopotamus gorgops Dietrich, 1928 | |
Hippopotamus gorgops is an extinct species of the genus Hippopotamus known from remains found in Northern Africa, Eastern Africa, and the Levant. One of the largest hippopotamus species, it first appeared during the late Pliocene-Early Pleistocene, and became extinct during the early Middle Pleistocene. [1]
The species was described by German scientist Wilhelm Otto Dietrich in 1928. [2] It is closely related and possibly ancestral to the large European species Hippopotamus antiquus . [3]
Hippopotamus gorgops is known from remains found across Northern and Eastern Africa, spanning from the Late Pliocene-Early Pleistocene boundary to the early Middle Pleistocene. [1] The species, along with other hippopotamids, was abundant at Buia in present-day Eritrea. [4] Remains assigned to H. gorgops alongside the species H. behemoth are known from the Ubeidiya locality in Israel, dating to approximately 1.6 million years ago, with some authors suggesting that H. behemoth is actually a synonym of H. gorgops, given their similar morphology. [5] [6]
H. gorgops grew substantially larger than the living hippopotamus (H. amphibius), [7] with an estimated body mass of over 4,000 kilograms (8,800 lb). [8] In comparison to H. amphibius, the orbits and nasal openings are elevated higher above the rest of the skull, with the temporal fossa being shorter, resulting in the distance between the orbits and occiput being shorter. This likely meant a more forward attachment of the temporal muscles. [1] The nuchal and sagittal crests were also proportionally taller, and the palate was elongate. [9] In life H. gorgops would have likely greatly resembled H. amphibius. [1]
Isotopic analysis of specimens from the Lake Turkana Basin suggests a high consumption of C4 plants. [10] For specimens from Olduvai Gorge in Tanzania, dental microwear suggests a mixed feeding diet (including both browse and grass), while mesowear suggests a largely grazing diet. [11] The elevated orbits have led to suggestions of a more aquatic lifestyle than H. amphibius, [12] though its robust limbs indicate that it was probably capable of moving efficiently on land, like H. amphibius. [9]
Remains with cut or fracture marks from sites across Africa from the Early Pleistocene indicate that H. gorgops was butchered by archaic humans. [13] [14] [15] These include El-Kherba in Algeria, dating to around 1.8 million years ago (mya), [15] Kilombe caldera in Kenya, dating to around 1.76 mya, [14] and the Buia site in Eritrea dating to the late Early Pleistocene, [13] around 1 mya. [9] Often at these sites tools from the Oldowan industry are present. [14] [15] At Kilombe and Buia, it is unclear whether the hippos were hunted or scavenged. [13] [14] It is unlikely that healthy adult individuals were hunted, as with living hippopotamuses, they were likely dangerous. [1]
Paranthropus is a genus of extinct hominin which contains two widely accepted species: P. robustus and P. boisei. However, the validity of Paranthropus is contested, and it is sometimes considered to be synonymous with Australopithecus. They are also referred to as the robust australopithecines. They lived between approximately 2.9 and 1.2 million years ago (mya) from the end of the Pliocene to the Middle Pleistocene.
The hippopotamus (; pl.: hippopotamuses; also shortened to hippo, further qualified as the common hippopotamus, Nile hippopotamus, or river hippopotamus, is a large semiaquatic mammal native to sub-Saharan Africa. It is one of only two extant species in the family Hippopotamidae, the other being the pygmy hippopotamus. Its name comes from the ancient Greek for "river horse".
Hippopotamidae is a family of stout, naked-skinned, and semiaquatic artiodactyl mammals, possessing three-chambered stomachs and walking on four toes on each foot. While they resemble pigs physiologically, their closest living relatives are the cetaceans. They are sometimes referred to as hippopotamids.
The pygmy hippopotamus or pygmy hippo is a small hippopotamid which is native to the forests and swamps of West Africa, primarily in Liberia, with small populations in Sierra Leone, Guinea, and Ivory Coast. It has been extirpated from Nigeria.
Hippopotamus is a genus of artiodactyl mammals consisting of one extant species, Hippopotamus amphibius, the river hippopotamus, and several extinct species from both recent and prehistoric times. It belongs to the family Hippopotamidae, which also includes the pygmy hippopotamus and a number of extinct genera.
Palaeoloxodon recki, often known by the synonym Elephas recki, is an extinct species of elephant native to Africa and West Asia from the Pliocene or Early Pleistocene to the Middle Pleistocene. During most of its existence, the species represented the dominant elephant species in East Africa. The species is divided into five roughly chronologically successive subspecies. While the type and latest subspecies P. recki recki as well as the preceding P. recki ileretensis are widely accepted to be closely related to Eurasian Palaeoloxodon, the relationships of the other, chronologically earlier subspecies to P. recki recki and P. recki ileretensis are uncertain, with it being suggested they are unrelated and should be elevated to separate species.
Equus namadicus is a prehistoric equid, known from remains dating to the Middle and Late Pleistocene from across the Indian subcontinent, with its last dated records being approximately 29-14,000 years ago. It is considered a "stenonine horse", meaning that it is probably more closely related to zebras and asses than true horses. It is relatively large in size. It is very similar to the earlier Equus sivalensis, also from the Indian subcontinent, from which it only differs in size and in subtle aspects of dental anatomy, and it has sometimes been suggested to be a synonym of it.
Hexaprotodon is an extinct genus of hippopotamid known from Asia and possibly Africa and Europe. The name Hexaprotodon means "six front teeth" as some of the fossil forms have three pairs of incisors. The pygmy hippopotamus was historically placed in the genus, but today is generally placed in its own genus. The core Asian members of the genus ranged from the Indian subcontinent to Southeast Asia, and are thought to have had an aquatic ecology similar to that of the living common hippopotamus. The last members of the genus became extinct during the Late Pleistocene, perhaps as recently as 15,000 years ago.
The Cypriot pygmy hippopotamus is an extinct species of dwarf hippopotamus that inhabited the island of Cyprus from the Pleistocene until the early Holocene. One the smallest known hippopotamus species, it was comparable in size to the living pygmy hippopotamus though it was more closely related to the common hippopotamus, with its small body size a result of insular dwarfism. It represented one of only two large terrestrial mammals on Cyprus alongside the Cyprus dwarf elephant. The species became extinct around 12,000 years ago following the arrival of humans on Cyprus, and potential evidence of human hunting has been found at the Aetokremnos rockshelter on the southern coast of the island.
Hippopotamus antiquus is an extinct species of the genus Hippopotamus that ranged across Europe during the Early and Middle Pleistocene. It was considerably larger than the living hippopotamus.
Hippopotamus melitensis is an extinct hippopotamus from Malta. It lived during Middle-Late Pleistocene. It probably descended from Hippopotamus pentlandi from Sicily, which in turn probably descended from the common hippopotamus. Like Hippopotamus pentlandi, Hippopotamus melitensis is substantially smaller than H. amphibius as a result of insular dwarfism, having an estimated mass of approximately 900 kg, which is smaller than the 1100 kg estimated for H. pentlandi. The diet of H. melitensis is suggested to have been more generalist than Hippopotamus amphibius, likely as a result of limited resource diversity and lack of competition, as the only other large herbivore on the island was the dwarf elephant Palaeoloxodon mnaidriensis. The majority of findings of this species are from Għar Dalam, a cave on Malta famous for its Pleistocene fossil deposits.
Hippopotamus pentlandi is an extinct species of hippopotamus from Sicily, known from the late Middle Pleistocene to early Late Pleistocene. It is the largest of the insular dwarf hippos known from the Pleistocene of the Mediterranean, "at most 20% smaller than the mainland forms", with an estimated body mass of approximately 1100 kg. It is suggested that it arrived in Sicily between 250,000 and 150,000 years ago, probably descending from the modern hippopotamus, with an origin from Hippopotamus antiquus being less likely. In comparison to those species, the muzzle was shorter, the occipital and nasal regions were more developed, the mastoid process was enlarged, and the dental row was shortened, and the condyle of the mandible is low. In comparison to H. amphibius, the orbits are also elevated. It was present in Sicily until at least the latest Middle Pleistocene around 120 kya, and was probably extinct by the beginning of Marine Isotope Stage 4. Contemporaneous species include the dwarf elephant Palaeoloxodon mnaidriensis, the aurochs, red deer, steppe bison, fallow deer, wild boar, brown bear, wolves, red foxes, cave hyena and cave lions. Its diet was likely grazing dominated, similar to that of modern H. amphibius. It is probably ancestral to Hippopotamus melitensis from Malta, which is substantially smaller than H. pentlandi.
Stephanorhinus is an extinct genus of two-horned rhinoceros native to Eurasia and North Africa that lived during the Late Pliocene to Late Pleistocene. Species of Stephanorhinus were the predominant and often only species of rhinoceros in much of temperate Eurasia, especially Europe, for most of the Pleistocene. The last two species of Stephanorhinus – Merck's rhinoceros and the narrow-nosed rhinoceros – went extinct during the last glacial period.
The narrow-nosed rhinoceros, also known as the steppe rhinoceros is an extinct species of rhinoceros belonging to the genus Stephanorhinus that lived in western Eurasia, including Europe, as well as North Africa during the Pleistocene. It first appeared in Europe around 500,000 years ago during the Middle Pleistocene and survived there until at least 34,000 years Before Present. It was native to temperate and Mediterranean environments, where it fed on low growing plants and to a lesser extent woody plants. Evidence has been found that it was exploited for food by archaic humans, including Neanderthals.
Several expansions of populations of archaic humans out of Africa and throughout Eurasia took place in the course of the Lower Paleolithic, and into the beginning Middle Paleolithic, between about 2.1 million and 0.2 million years ago (Ma). These expansions are collectively known as Out of Africa I, in contrast to the expansion of Homo sapiens (anatomically modern humans) into Eurasia, which may have begun shortly after 0.2 million years ago.
Hippopotamus behemoth is an extinct species of hippopotamus from the Early Pleistocene of the Levant. Fossils of it, and its probable ancestor, H. gorgops, are found in the ‘Ubeidiya site in the southern Levant, dating to the Early Pleistocene, around 1.4 million years ago.
"Crocodylus" gariepensis is an extinct species of crocodile that lived in southern Africa during the Early Miocene about 17.5 million years ago (Ma). Fossils have been found along a bank of the Orange River in Namibia, near its border with South Africa.
Buya or Buia is an archaeological site in the Danakil Depression of Eritrea. It is known for the discovery of Madam Buya, a one million-year-old fossil of a Homo erectus skull. Two other expeditions in 2011 and 2012 also unearthed ancient hominid fossils at the site. Archaeologists have uncovered large quantities of animal fossils and lithic tools in the area.
Pliorhinus is an extinct genus of rhinoceros known from the Late Miocene and Pliocene of Eurasia. The type species, Pliorhinus megarhinus, was previously assigned to Dihoplus.
Bos buiaensis is an extinct species of cattle. The species is known from a million year old skull fossil found at the archaeological site of Buya, Eritrea in 2003. It was reassembled by excavators from over one hundred shards.