Ictidorhinus Temporal range: Late Permian | |
---|---|
Life restoration of Ictidorhinus martinsi | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Suborder: | † Biarmosuchia |
Family: | † Ictidorhinidae Broom, 1932 |
Genus: | † Ictidorhinus Broom, 1913 |
Type species | |
†Ictidorhinus martinsi Broom, 1913 |
Ictidorhinus is an extinct genus of biarmosuchian therapsids. [1] Fossils have been found from the Dicynodon Assemblage Zone of the Beaufort Group in the Karoo Basin, South Africa and are of Late Permian age. It had a short snout and proportionally large orbits. These characteristics may be representative of a juvenile animal, possibly of Lycaenodon . However, these two genera are not known to have existed at the same time, making it unlikely for Ictidorhinus material to be from a juvenile form of Lycaenodon. [2]
Ictidorhinus is the best-known representative of the family Ictidorhinidae, named by South African paleontologist Robert Broom in 1932. Many biarmosuchians have been identified as ictidorhinids since the family was first named, mostly on the basis of their small size. Several biarmosuchians from Russia have been classified as ictidorhinids on the basis of partial jaw bones alone. Recent phylogenetic studies of biarmosuchians have found that Ictidorhinidae is a paraphyletic collection of species that fall outside the more derived clade Burnetiamorpha. [3]
Therapsida is a clade comprising a major group of eupelycosaurian synapsids that includes mammals and their ancestors and close relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, resulting in a more "standing" quadrupedal posture, as opposed to the lower sprawling posture of many reptiles and amphibians.
Dinocephalians are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania.
Eotitanosuchus is an extinct genus of biarmosuchian therapsids whose fossils were found in the town of Ochyor in Perm Krai, Russia. It lived about 267 million years ago. The only species is Eotitanosuchus olsoni.
Biarmosuchia is an extinct clade of non-mammalian synapsids from the Permian. Biarmosuchians are the most basal group of the therapsids. They were moderately-sized, lightly built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation.
The Tapinocephalus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the middle Abrahamskraal Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching approximately 2,000 metres (6,600 ft), occur from Merweville and Leeu-Gamka in its southernmost exposures, from Sutherland through to Beaufort West where outcrops start to only be found in the south-east, north of Oudshoorn and Willowmore, reaching up to areas south of Graaff-Reinet. Its northernmost exposures occur around the towns Fraserburg and Victoria West. The Tapinocephalus Assemblage Zone is the second biozone of the Beaufort Group.
Burnetiidae is an extinct family of biarmosuchian therapsids that lived in the Permian period whose fossils are found in South Africa, Zambia and Russia. It contains Bondoceras, Bullacephalus, Burnetia, Mobaceras, Niuksenitia, Paraburnetia and Proburnetia.
Lemurosaurus is a genus of extinct biarmosuchian therapsids from the Late Permian of South Africa. The generic epithet Lemursaurus is a mix of Latin, lemures “ghosts, spirits”, and Greek, sauros, “lizard”. Lemurosaurus is easily identifiable by its prominent eye crests, and large eyes. The name Lemurosaurus pricei was coined by paleontologist Robert Broom in 1949, based on a single small crushed skull, measured at approximately 86 millimeters in length, found on the Dorsfontein farm in Graaff-Reinet. To date, only two skulls of the Lemurosaurus have been discovered, so body size is unknown. The second larger, more intact, skull was found in 1974 by a team from the National Museum, Bloemfontein.
Burnetia is an extinct genus of biarmosuchian therapsids in the family Burnetiidae, from the Late Permian of South Africa. Burnetia is known so far from a single holotype skull lacking the lower jaws described by South African paleontologist Robert Broom in 1923. Due to erosion and dorsoventral crushing, features of the skull are hard to interpret. Stutural lines are further distorted by the unusual shape of the skull roof, including many bosses and protuberances.
Paraburnetia is an extinct genus of biarmosuchian therapsids from the Late Permian of South Africa. It is known for its species P. sneeubergensis and belongs to the family Burnetiidae. Paraburnetia lived just before the Permian–Triassic mass extinction event.
Lobalopex is an extinct genus of biarmosuchian therapsids. It was alive during the Late Permian and has only been found in the Teekloof Formation in South Africa. The only known species of the genus is Lobalopex mordax. Lobalopex is part of the clade of Burnetiamorpha, which have fossil specimens located in multiple areas of Africa and Russia.
Bullacephalus is an extinct genus of biarmosuchian therapsids belonging to the family Burnetiidae. The type species B. jacksoni was named in 2003. It is known from a relatively complete skull and lower jaw, discovered in the Late Permian Tapinocephalus Assemblage Zone of the Beaufort Group of South Africa. This genus of therapsida lived during the Late Permian period, approximately 250 million years ago.
Hipposauroides is an extinct genus of biarmosuchian therapsids from the Late Permian of South Africa.
Lycaenodon is an extinct genus of biarmosuchian therapsids from the Late Permian of South Africa. It is known from a single species, Lycaenodon longiceps, which was named by South African paleontologist Robert Broom in 1925. Both are small-bodied biarmosuchians. Two specimens are known, and both preserve only the front portions of the skull. These specimens come from the Cistecephalus Assemblage Zone of the Karoo Basin. Broom attributed the back portion of a third skull to Lycaenodon, but subsequent examiners considered it to belong to a gorgonopsian or dinocephalian and not a biarmosuchian. Most of the distinguishing features of Lycaenodon come from its palate. As a member of Biarmosuchia, the most basal group of therapsids, Lycaenodon shares many features with earlier and less mammal-like synapsids like Dimetrodon.
Niaftasuchus is an extinct genus of therapsids. Its type and only named species is Niaftasuchus zekkeli.
Pachydectes is an extinct genus of biarmosuchian therapsids from the Middle Permian of South Africa known from a single skull. The etymology of the name Pachydectes is derived from the Greek word pakhus, meaning "thick" or "thickened", and dektes, meaning "biter". In conjunction this name is representative of the unique pachyostotic bone present above the maxillary canine tooth found in the skull of the specimen. There is only one known species within the genus, Pachydectes elsi which is named in honor of the person who discovered the fossil.
Herpetoskylax is an extinct genus of biarmosuchian therapsids which existed in South Africa. The type species is Herpetoskylax hopsoni. It lived in the Late Permian Period.
Hipposauridae is an extinct family of biarmosuchian therapsids from the Late Permian of South Africa. It includes the genus Hipposaurus, and possibly the genera Hipposauroides and Pseudhipposaurus. A closely related biarmosuchian is Ictidorhinus, which has been placed in the family Ictidorhinidae.
Ustia is an extinct genus of biarmosuchian therapsids from the Middle Permian of Russia. It is known from a single species, Ustia atra, which was described in 2003 from an isolated lower jaw. Ustia was classified in the family Ictidorhinidae, which also includes the genus Ictidorhinus from South Africa. Both are relatively small biarmosuchians. Several other Russian therapsids known only from lower jaw bones have been placed in Ictidorhinidae, and the family is likely a paraphyletic assemblage representing a small body type than a true clade.
Rubidgina is a genus of Biarmosuchian therapsid from Patrysfontein, Wellwood, South Africa known from RC 55, a skull with lower jaws. This specimen is a putative juvenile. It has been suggested that this specimen actually represents a juvenile of Herpetoskylax hopsoni. However, because the specimen lacks distinctive features, it cannot be determined if it is actually a juvenile of Herpetoskylax or if its current name of Rubidgina should remain.
Impumlophantsi is an extinct genus of biarmosuchian therapsids from the Permian Abrahamskraal Formation of South Africa. The genus contains a single species, I. boonstrai, known from a partial skull and fragmentary postcrania.