Lactarius helvus | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Fungi |
Division: | Basidiomycota |
Class: | Agaricomycetes |
Order: | Russulales |
Family: | Russulaceae |
Genus: | Lactarius |
Species: | L. helvus |
Binomial name | |
Lactarius helvus (Fr.) Fr. (1838) | |
Synonyms | |
Lactarius helvus | |
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Gills on hymenium | |
Cap is depressed | |
Hymenium is decurrent | |
Stipe is bare | |
Spore print is tan | |
Ecology is mycorrhizal | |
Edibility is poisonous |
Lactarius helvus, commonly known as fenugreek milkcap, is a member of the large milkcap genus Lactarius in the order Russulales. Fruiting bodies can be found in Sphagnum moss in coniferous and deciduous woodland in Europe, and possibly North America, although considerable debate continues about the North American variety, formerly referred to as Lactarius aquifluus. Mushrooms are pale brown-grey or beige in colour and funnel-shaped, with colourless, watery milk. Its distinctive smell has been likened to fenugreek, celery, liquorice, or Maggi instant soup. Mildly toxic when raw, it has been implicated in the poisoning of 418 people near Leipzig in October 1949. However, it is used in small quantities as a spice when dried. Sotolon, the agent that gives the fungus its odour, also occurs in fenugreek, maple syrup and lovage.
The possible North American variety has a more salmon to pink color with scant watery milk and a strong smell of maple syrup or caramel. In Quebec it is known as the maple milky cap (Lactaire à odeur d'érable), and is fairly commonly eaten cooked and also sold as a dried powder and used as a spice, much in the same way that the candy cap mushroom is used on the west coast. Thus far, there have been no reported cases of illness from this variety in North America. Some confusion with the species Lactarius quietus var incanus is possible due to the similar smell. Lactarius quietus var incus, however, has a darker and more red appearance and when young, its milk is opaque and white, as opposed to the clear milk in Lactarius helvus. Furthermore, the possible North American variety of Lactarius helvus is restricted in distribution to the great lakes and north east, while lactarius quietus var incanus is commonly found much further south.
It was initially described by Elias Magnus Fries in 1821 as Agaricus helvus, before he placed it in the genus Lactarius in 1838. Peck's Lactarius aquifluus has been deemed a synonym. The specific epithet helvus is derived from the Latin for 'honey-yellow'. Its name in German is Maggipilz. S
The velvety cap is initially slightly convex (planoconvex), becoming funnel-shaped (infundibuliform) as it matures; it is 4–15 cm (1½–4 in) in diameter and has a faint zonate (bull's-eye) pattern, beige or light grey at the margins and darkening toward the centre. The decurrent gills are cream when young, and darkening to ochre-yellow with age. The flesh is white or beige, often pink-tinged. The latex, or milk, is watery and colourless, unlike that of any other milkcap[ citation needed ]. The hollow cylindrical stalk is 3–9 cm (1–3½ in) high by 1–2 cm (½ in) wide. It may be whitish when young, ochre or pink-tinged, and sometimes spotted orange-brown. The highly distinctive smell has been described as reminiscent of fenugreek, with a mild taste. [1] To others, the smell has been likened to lovage or celery, [2] or liquorice or a packet of Maggi instant soup. [3]
Fruiting bodies are found in groups in woodlands near conifers such as Picea and Pinus , as well as the deciduous birch ( Betula ) and rarely beech ( Fagus ). More specifically, they occur in wet places, often growing in Sphagnum , in late summer and autumn. The species is found across Europe and Asia. [1]
Symptoms of poisoning are reported to occur within thirty minutes of consumption, with nausea and vomiting accompanied by vertigo and chills. The toxic agents are thought to be sesquiterpenes. In October 1949, 418 people were poisoned near Leipzig in eastern Germany. [4] The toxins are destroyed by thorough boiling, and the species is used in small quantities as a spice after drying. [3]
The agent which gives the fungus its distinctive odour is sotolon, [5] which also gives fenugreek seed and lovage their characteristic smells. [6] It is present as well in molasses, aged sake and white wine, flor sherry, and roasted tobacco, [7] as well as in maple syrup. [8]
Edible mushrooms are the fleshy fruit bodies of numerous species of macrofungi. Edibility may be defined by criteria including the absence of poisonous effects on humans and desirable taste and aroma. Mushrooms that have a particularly desirable taste are described as "choice". Edible mushrooms are consumed for their nutritional and culinary value. Mushrooms, especially dried shiitake, are sources of umami flavor.
Fenugreek is an annual plant in the family Fabaceae, with leaves consisting of three small obovate to oblong leaflets. It is cultivated worldwide as a semiarid crop. Its leaves and seeds are common ingredients in dishes from the Indian subcontinent, and have been used as a culinary ingredient since ancient times. Its use as a food ingredient in small quantities is safe.
The Russulaceae are a diverse family of fungi in the order Russulales, with roughly 1,900 known species and a worldwide distribution. They comprise the brittlegills and the milk-caps, well-known mushroom-forming fungi that include some edible species. These gilled mushrooms are characterised by the brittle flesh of their fruitbodies.
Sotolon is a butenolide lactone and an extremely potent aroma compound, with the typical smell of fenugreek or curry at high concentrations and maple syrup, caramel, or burnt sugar at lower concentrations. Sotolon is the major aroma and flavor component of fenugreek seed and lovage, and is one of several aromatic and flavor components of artificial maple syrup. It is also present in molasses, aged rum, aged sake and white wine, flor sherry, roast tobacco, and dried fruiting bodies of the mushroom Lactarius helvus. Sotolon can pass through the body relatively unchanged, and consumption of foods high in sotolon, such as fenugreek, can impart a maple syrup aroma to one's sweat and urine. In some individuals with the genetic disorder maple syrup urine disease, sotolon is spontaneously produced in their bodies and excreted in their urine, leading to the disease's characteristic smell.
Lactarius is a genus of mushroom-producing, ectomycorrhizal fungi, containing several edible species. The species of the genus, commonly known as milk-caps, are characterized by the milky fluid ("latex") they exude when cut or damaged. Like the closely related genus Russula, their flesh has a distinctive brittle consistency. It is a large genus with over 500 known species, mainly distributed in the Northern hemisphere. Recently, the genus Lactifluus has been separated from Lactarius based on molecular phylogenetic evidence.
Lactarius torminosus, commonly known as the woolly milkcap or the bearded milkcap, is a large species of agaric fungus. A common and widely distributed species, it is found in North Africa, northern Asia, Europe, and North America. It was first described scientifically by Jacob Christian Schäffer in 1774 as an Agaricus, and later transferred to the genus Lactarius in 1821 by Samuel Frederick Gray. A variety, L. torminosus var. nordmanensis, is known from the United States, Canada, and Switzerland. L. torminosus officially became the type species of Lactarius in 2011 after molecular studies prompted the taxonomic reshuffling of species between several Russulaceae genera.
Candy cap or curry milkcap is the English-language common name for two closely related edible species of Lactarius: Lactarius camphoratus and Lactarius rubidus. Additionally, L. rufulus is termed the southern candy cap. Many similar species are known.
Lactarius uvidus, commonly known as the purple staining milk cap, is a European and North American "milk-cap" mushroom, of which the milk turns violet when the flesh is damaged. The fungi generally identified as L. uvidus are part of a complex of closely related species and varieties which are difficult to delimit definitively.
Lactifluus piperatus, commonly known as the blancaccio, is a basidiomycete fungus of the genus Lactifluus. It was the original type species of the genus Lactarius, before being moved to Lactifluus.
Lactarius blennius is a medium-sized mushroom of the genus Lactarius found commonly in beech forests in Europe, where it is mycorrhizal, favouring the European beech. It was first described by Elias Magnus Fries. Though its colour and size vary, it is distinctive because it is slimy when wet and exudes copious amounts of milk. It has been the subject of some chemical research, and it can be used to produce pigments and blennins. Blennins, some of which have shown potential medical application, are derived from lactarane, a chemical so named because of their association with Lactarius. The edibility of L. blennius is uncertain, with different mycologists suggesting that it is edible, inedible or even poisonous.
Lactifluus volemus, formerly known as Lactarius volemus, and commonly known as the weeping milk cap or bradley, is a species of fungus in the family Russulaceae. Phylogenetic analysis suggests that L. volemus represents several species or subspecies, rather than a single taxon.
Lactarius tabidus, commonly known as the birch milkcap, is an inedible mushroom of the genus Lactarius. It can be found in North America and Europe, and grows at the base of pine in Autumn. Its white milk stains fabric a sulphur yellow, from which its specific epithet derives.
Lactarius vietus is a species of fungus in the family Russulaceae, first described by Elias Magnus Fries. It produces moderately sized and brittle mushrooms, which grow on the forest floor or on rotting wood. The flattened-convex cap can vary in shape, sometimes forming the shape of a wide funnel. It is typically grey, but the colour varies. The species has crowded, light-coloured gills, which produce white milk. The spore print is typically whitish, but also varies considerably. The mushrooms typically have a strong, acrid taste and have been described as inedible, but other authors have described them as consumable after boiling. L. vietus feeds by forming an ectomycorrhizal relationship with surrounding trees, and it favours birch. It grows in autumn months and is fairly common in Europe, North America and eastern Asia.
Lactarius quietus is a mushroom of the genus Lactarius. It is easily identified by its oily scent and the concentric bands on its cap. It is brown, and is probably named after its matte, "quiet" surface and colouration. It is found exclusively under oak trees in Europe, where it grows solitarily or in scattered groups in autumn months. In North America, the variety L. quietus var. incanus is fairly common in the same habitat. The mushroom's edibility is disputed.
Lactarius alnicola, commonly known as the golden milkcap, is a species of fungus in the family Russulaceae. The fruit bodies produced by the fungus are characterized by a sticky, vanilla-colored cap up to 20 cm (8 in) wide with a mixture of yellow tones arranged in faint concentric bands. The stem is up to 5 cm (2 in) long and has yellow-brown spots. When it is cut or injured, the mushroom oozes a white latex, which has an intensely peppery taste. The acrid taste of the fruit bodies renders them unpalatable. Two varieties have been named: var. pitkinensis, known from Colorado, and var. pungens, from Michigan.
Lactarius repraesentaneus, commonly known as the northern bearded milkcap, the northern milkcap, or the purple-staining bearded milkcap, is a species of fungus in the family Russulaceae. It has a northerly distribution, and is found in temperate regions of North America and Europe, associated with spruce trees. Distinguishing features of its fruit body include the large orange-yellow cap up to 18 cm (7.1 in) wide, cream to pale yellow gills, and a yellow coarsely-pitted stem that is up to 12 cm (4.7 in) long and 3 cm (1.2 in) thick. Cut fruit bodies ooze a white latex that will stain mushroom tissue lilac to purple. Several chemicals have been isolated and identified from the fruit bodies that can modify the growth of plants, and the mushroom also has antibiotic activity against Staphylococcus aureus. L. repraesentaneus is poisonous, and consumption causes stomach aches.
Lactifluus deceptivus, commonly known as the deceiving milkcap, is a common species of fungus in the family Russulaceae.
Lactarius vinaceorufescens, commonly known as the yellow-staining milkcap or the yellow-latex milky, is a poisonous species of fungus in the family Russulaceae. It produces mushrooms with pinkish-cinnamon caps up to 12 cm (4.7 in) wide held by pinkish-white stems up to 7 cm (2.8 in) long. The closely spaced whitish to pinkish buff gills develop wine-red spots in age. When it is cut or injured, the mushroom oozes a white latex that rapidly turns bright sulfur-yellow. The species, common and widely distributed in North America, grows in the ground in association with conifer trees. There are several other Lactarius species that bear resemblance to L. vinaceorufescens, but most can be distinguished by differences in staining reactions, macroscopic characteristics, or habitat.
Lactifluus clarkeae, formerly known as Lactarius clarkeae, is a species of mushroom-forming fungus in the family Russulaceae. It is found in Australia and New Zealand in mycorrhizal association with species of Nothofagus and the family Myrtaceae.
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