List of lichens named by Carl Linnaeus

The Swedish taxonomist Carl Linnaeus described 110 lichen species in his various works between 1753 and 1774, chiefly in the first (1753) and second (1763) editions of Species Plantarum , and in later works such as Systema Naturae and Mantissa plantarum. Although Linnaeus showed much less interest in lichens than in flowering plants, he recognized their ecological role as pioneer organisms in primary ecological successions. Contrary to popular belief, ^{[1] [2] [3]} he never referred to lichens as the rustici pauperrimi ("the poorest peasants") or "poor trash of nature": that phrase was introduced by Georg Franz Hoffmann in 1787 and later misattributed to Linnaeus himself. ^[4]

Of the roughly 2,000 lichen species now known from northern Europe, Linnaeus recognized only about 5% in his treatments. Most of his lichen names were based on material from northern Europe, especially Sweden, with only a handful of species described from specimens collected outside Europe. His lichen accounts were usually brief, consisting of short diagnostic phrases with few details of morphology, variation or ecology when judged against modern species descriptions. Even so, the Linnaean Herbarium at the Linnean Society of London contains 324 sheets with lichens and lichen-like organisms, although not all of these represent original material available to him at the time of publication. ^[5]

Many of Linnaeus's lichen names are still in use today, though now placed in genera that did not exist in his time, and his species serve as the types for several important lichen genera including Cladonia , Lecidea , Lobaria , Parmelia , Peltigera , Ramalina and Usnea . ^[5] At the same time, some of his species concepts were broad by modern standards. For example, his Lichen subfuscus and Lichen geographicus each correspond to species complexes that are now divided into several segregate taxa on morphological and molecular grounds. ^[6] Other Linnaean names have been formally rejected or conserved with different types to preserve prevailing usage and stabilize nomenclature. ^[5]

The typification and interpretation of Linnaean lichen names has therefore required sustained effort by lichenologists, involving critical study of his specimens, publications and other historical sources to determine the correct application of each name under the International Code of Nomenclature for algae, fungi, and plants. ^[7] This list assembles all 110 Linnaean names that Linnaeus intended for lichens in the modern sense, regardless of their present status. For each original Linnaean name (the basionym), it records the original year of publication, the current taxonomic status or accepted name, and the year in which the present combination was published. In the "Basionym" column, author citations are omitted because all names in the list were originally published by Linnaeus (abbreviated "L."). The treatment largely follows the typifications and interpretations of Per Magnus Jørgensen, Peter Wilfred James and Charles Edward Jarvis published in 1994, supplemented by later taxonomic revisions and by current usage in major nomenclatural databases such as MycoBank and Index Fungorum.

Lichen taxa described by Carl Linnaeus and their current names

Basionym	Year of publication	Current name / status	Authority	Year of recombination	Image
Byssus botryoides [8]	1753	Lichenomphalia umbellifera	(L.) Redhead, Lutzoni, Moncalvo & Vilgalys [9]	2008
Byssus candelaris [8]	1753	Chrysothrix candelaris	(L.) J.R.Laundon [10]	1981
Byssus incana [8]	1753	Lepraria incana	(L.) Ach. [11]	1803
Byssus lacteus [8]	1753	Nom. rej. [note 1]	N/A	N/A	N/A
Lichen ampullaceus [18]	1753	Parmelia omphalodes [note 2]	(L.) Ach.	1803	N/A
Lichen aphtosus [23]	1753	Peltigera aphthosa	(L.) Willd. [24]	1787
Lichen aquaticus [23]	1753	Dermatocarpon luridum	(Dill. ex With.) J.R.Laundon [25]	1984
Lichen arcticus [23]	1753	Nephroma arcticum	(L.) Torss. [26]	1843
Lichen articulatus [27]	1753	Usnea articulata	(L.) Hoffm. [28]	1796
Lichen atro-albus [29]	1753	Nom. rej. [note 3]	N/A	N/A	N/A
Lichen atro-virens [29]	1753	Nom. rej. [note 4]	N/A	N/A	N/A
Lichen barbatus [33]	1753	Usnea barbata	(L.) F.H.Wigg.	1780
Lichen burgessii [34]	1774	Leptogium burgessii	(L.) Mont. [35]	1840
Lichen byssoides [36]	1767	Baeomyces rufus	(L.) P.Gaertn., B.Mey. & Scherb.	1802
Lichen calcareus [37]	1753	Circinaria calcarea	(L.) A.Nordin, Savić & Tibell [38]	2010
Lichen calicaris [18]	1753	Ramalina calicaris	(L.) Röhl. [39]	1813
Lichen candelarius [29]	1753	Polycauliona candelaria	(L.) Frödén, Arup & Søchting [40]	2013
Lichen caninus [41]	1753	Peltigera canina	(L.) Willd. [24]	1787
Lichen caperatus [42]	1753	Flavoparmelia caperata	(L.) Hale [43]	1986
Lichen carpineus [29]	1753	Glaucomaria carpinea	(L.) S.Y.Kondr., L.Lőkös & Farkas [44]	2019
Lichen centrifugus [45]	1753	Arctoparmelia centrifuga	(L.) Hale [46]	1986
Lichen chalybeiformis [33]	1753	Bryoria capillaris	(L.) Brodo & D.Hawksw. [47]	1977
Lichen chrysophthalmus [48]	1771	Teloschistes chrysophthalmus	(L.) Th.Fr.	1861
Lichen ciliaris [49]	1753	Anaptychia ciliaris	(L.) Körb. ex A.Massal.	1853
Lichen cinereus [50]	1767	Aspicilia cinerea	(L.) Körb.	1855
Lichen cocciferus [51]	1753	Cladonia coccifera	(L.) Willd.	1787
Lichen corallinus [52]	1767	Lepra corallina	(L.) Hafellner	2016
Lichen cornucopioides [51]	1753	Nom. rej. [note 5]	N/A	N/A	N/A
Lichen cornutus [55]	1753	Cladonia cornuta	(L.) Hoffm.	1791
Lichen cristatus [56]	1753	Lathagrium cristatum	(L.) Otálora, P.M.Jørg. & Wedin	2014
Lichen crocatus [57]	1771	Pseudocyphellaria crocata	(L.) Vain.	1898
Lichen croceus [41]	1753	Solorina crocea	(L.) Ach.	1808
Lichen cylindricus [49]	1753	Umbilicaria cylindrica	(L.) Delise	1830
Lichen deformis [55]	1753	Cladonia deformis	(L.) Hoffm.	1796
Lichen deustus [58]	1753	Umbilicaria deusta	(L.) Baumg.	1790
Lichen digitatus [55]	1753	Cladonia digitata	(L.) Hoffm.	1796
Lichen divaricatus [59]	1767	Evernia divaricata	(L.) Ach.	1810
Lichen ericetorum [29]	1753	Icmadophila ericetorum	(L.) Zahlbr.	1895
Lichen fagineus [29]	1753	Lepra amara	(Ach.) Hafellner	2016
Lichen fahlunensis [56]	1753	Nom. rej. [note 6]	style="text-align:center;" | N/A	N/A	N/A
Lichen farinaceus [18]	1753	Ramalina farinacea	(L.) Ach.	1810
Lichen fascicularis [36]	1767	Gabura fascicularis	(L.) Otálora & Wedin [62]	2013
Lichen fimbriatus [55]	1753	Cladonia fimbriata	(L.) Fr.	1831
Lichen floridus [27]	1753	Usnea florida	(L.) F.H.Wigg.	1780
Lichen fragilis [63]	1753	Sphaerophorus fragilis	(L.) Pers.	1794
Lichen fraxineus [18]	1753	Ramalina fraxinea	(L.) Ach.	1810
Lichen fuciformis [42]	1753	Roccella fuciformis	(L.) DC.	1805
Lichen furfuraceus [18]	1753	Pseudevernia furfuracea var. furfuracea	(L.) Zopf	1903
Lichen fusco-ater [37]	1753	Lecidea fuscoatra	(L.) Ach.	1803
Lichen gelidus [36]	1767	Placopsis gelida	(L.) Linds.	1866
Lichen geographicus [37]	1753	Rhizocarpon geographicum	(L.) DC.	1805
Lichen glaucus [23]	1753	Platismatia glauca	(L.) W.L.Culb. & C.F.Culb. [64]	1968
Lichen globiferus [36]	1767	Sphaerophorus globosus	(Huds.) Vain.	1903
Lichen gracilis [55]	1753	Cladonia gracilis	(L.) Willd.	1787
Lichen hirtus [33]	1753	Usnea hirta	(L.) F.H.Wigg.	1780
Lichen islandicus [65]	1753	Cetraria islandica subsp. islandica	(L.) Ach.	1803
Lichen islandicus var. tenuissimus [65]	1753	Cetraria aculeata	(Schreb.) Fr.	1826
Lichen jubatus [33]	1753	Nom. rej. [note 7]	N/A	N/A	N/A
Lichen juniperinus [42]	1753	Vulpicida juniperinus	(L.) J.-E.Mattsson & M.J.Lai	1993
Lichen lacteus [50]	1767	Varicellaria lactea	(L.) I.Schmitt & Lumbsch	2012
Lichen lanatus [33]	1753	Ephebe lanata	(L.) Vain.	1888
Lichen leucomelos [70]	1763	Leucodermia leucomelos	(L.) Kalb	2015
Lichen miniatus [41]	1753	Dermatocarpon miniatum var. miniatum	(L.) W.Mann	1825
Lichen nivalis [41]	1753	Nephromopsis nivalis	(L.) Divakar, A.Crespo & Lumbsch	2017
Lichen olivaceus [56]	1753	Melanohalea olivacea	(L.) O.Blanco, A.Crespo, Divakar, Essl., D.Hawksw. & Lumbsch	2004
Lichen omphalodes [56]	1753	Parmelia omphalodes	(L.) Ach.	1803
Lichen pallescens [45]	1753	Ochrolechia pallescens	(L.) A.Massal.	1853
Lichen parellus [50]	1767	Ochrolechia parella f. parella	(L.) A.Massal.	1852
Lichen parietinus [56]	1753	Xanthoria parietina var. parietina	(L.) Th.Fr.	1860
Lichen paschalis [71]	1753	Stereocaulon paschale	(L.) Hoffm.	1796
Lichen pertusus [72] Nom. cons. prop. [73]	1767	Pertusaria pertusa	(L.) Tuck.	1845
Lichen physodes [49]	1753	Hypogymnia physodes	(L.) Nyl.	1896
Lichen plicatus [63]	1753	Nom. rej. [note 8]	N/A	N/A	N/A
Lichen polyphyllus [58]	1753	Umbilicaria polyphylla	(L.) Baumg.	1790
Lichen polyrhizos [58]	1753	Umbilicaria polyrrhiza	(L.) Fr.	1825
Lichen proboscideus [58]	1753	Umbilicaria proboscidea	(L.) Schrad.	1794
Lichen prunastri [42]	1753	Evernia prunastri	(L.) Ach.	1810
Lichen pubescens [33]	1753	Pseudephebe pubescens	(L.) M.Choisy	1930
Lichen pulmonarius [65]	1753	Lobaria pulmonaria	(L.) Hoffm. [77]	1796
Lichen pustulatus [58]	1753	Lasallia pustulata	(L.) Mérat	1821
Lichen pyxidatus [51]	1753	Cladonia pyxidata	(L.) Hoffm.	1796
Lichen rangiferinus [71]	1753	Cladonia rangiferina	(L.) Weber ex F.H.Wigg.	1780
Lichen rangiferinus var. alpestris [71]	1753	Cladonia stellaris	(Opiz) Pouzar & Vězda	1971
Lichen rangiferinus var. sylvaticus [71]	1753	Cladonia portentosa	(Dufour) Coem.	1865
Lichen resupinatus [23]	1753	Nephroma resupinatum	(L.) Ach. [78]	1810
Lichen roccella [63]	1753	Roccella tinctoria [note 9]	DC.	1805
Lichen rugosus [37]	1753	Ascodichaena rugosa	Butin [82]	1977
Lichen rupicola [50]	1767	Lecanora rupicola var. rupicola	(L.) Zahlbr.	1928
Lichen saccatus [83]	1755	Solorina saccata	(L.) Ach.	1808
Lichen sanguinarius [37]	1753	Mycoblastus sanguinarius f. sanguinarius	(L.) Norman [84]	1852
Lichen saxatilis [45]	1753	Parmelia saxatilis	(L.) Ach. [11]	1803
Lichen scriptus [37]	1753	Graphis scripta	(L.) Ach.	1809
Lichen stellaris [49]	1753	Physcia stellaris	(L.) Nyl.	1853
Lichen stygius [56]	1753	Melanelia stygia	(L.) Essl. [85]	1978
Lichen subfuscus [45]	1753	Nom. rej. [note 10]	N/A	N/A	N/A
Lichen subulatus [71]	1753	Cladonia subulata	(L.) Weber ex F.H.Wigg.	1780
Lichen tartareus [29]	1753	Ochrolechia tartarea	(L.) A.Massal.	1852
Lichen uncialis [71]	1753	Cladonia uncialis subsp. uncialis	(L.) Weber ex F.H.Wigg.	1780
Lichen upsaliensis [45]	1753	Ochrolechia upsaliensis	(L.) A.Massal.	1852
Lichen usnea [52]	1767	Ramalina usnea	(L.) R.Howe	1914
Lichen velleus [58]	1753	Umbilicaria vellea	(L.) Ach.	1794
Lichen venosus [23]	1753	Peltigera venosa	(L.) Hoffm.	1789
Lichen ventosus [29]	1753	Ophioparma ventosa	(L.) Norman	1852
Lichen vernalis [90]	1768	Biatora vernalis	(L.) Fr.	1822
Lichen vulpinus [33]	1753	Letharia vulpina	(L.) Hue [91]	1899
Mucor fulvus [92]	1753	Chaenotheca furfuracea [note 11]	(L.) Tibell	1984
Mucor furfuraceus [92]	1753	Chaenotheca furfuracea	(L.) Tibell [96]	1984
Mucor lichenoides [92]	1753	Calicium salicinum [note 12]	Pers. [99]	1794
Mucor sphaerocephalus [92]	1753	Calicium lenticulare [note 13]	Ach.	1816	N/A
Tremella lichenoides [101]	1753	Scytinium lichenoides	(L.) Otálora, P.M.Jørg. & Wedin [102]	2014

Glossary

• Art. 56 – the article of the International Code of Nomenclature for algae, fungi, and plants (ICNafp) that allows a scientific name to be formally rejected when its continued use would cause confusion or instability (for example, because the original description is unidentifiable, ambiguous, or threatens a well-established name). A name rejected under Art. 56 becomes unavailable for use and is listed as nom. rej.
• basionym – the original name on which a new name is based
• nom. cons. prop. – nomen conservandum propositum; meaning "a name proposed for conservation". It indicates that a formal proposal has been submitted to conserve a particular name against an earlier competing one under Article 14 of the ICNafp
• nom. illeg. – nomen illegitimum; an illegitimate name according to the ICNafp
• nom. rej. – nomen rejiciendum; a name that has been formally rejected and is no longer available for use
• nom. rej. prop. – nomen rejiciendum propositum; as above, but in the proposal stage and not yet ratified

Notes

1. Byssus lacteus L. was introduced in Species plantarum (1753) from a brief diagnosis and a reference to Dillenius's figure in Historia muscorum. ^[12] Jørgensen, James and Jarvis found no original material in the Linnaean Herbarium, rejected an earlier typification, and showed that the associated Dillenian material in Oxford represents two species of Ochrolechia that match neither Linnaeus's diagnosis nor the illustration. ^[13] Because the original material cannot be tied with confidence to any species, they treated Byssus lacteus as a species non satis nota and proposed its rejection under Art. 56 of the Code. ^{[14] [15]} The Nomenclature Committee for Fungi accepted this proposal, and Index Fungorum now lists Byssus lacteus as a rejected name (nom. rej.) based on the Dillenian figure as holotype, placing it in Trentepohliaceae (Chlorophyta). ^{[16] [17]} The name is not used in lichen taxonomy and has sometimes been misapplied to Lichen lacteus L. (= Pertusaria lactea (L.) F.Arnold). ^{[13] [15]}
2. Lichen ampullaceus L. was described in Species plantarum (1753) from a short diagnosis that cited an illustration in Dillenius's Historia muscorum and a work by van Royen, and Linnaeus appears to have had no material of his own. ^{[19] [20]} No corresponding specimen exists in the Linnaean Herbarium, and only an inserted drawing occurs in the Dillenian herbarium. The Dillenian figure was long interpreted as showing monstrous forms of what is now Platismatia glauca infected by parasites, and on that basis Culberson & Culberson treated L. ampullaceus as a rejectable monstrosity under the provisions of the Code then in force, a procedure no longer permitted. Re-examining Dillenius's text and illustration, and a later colour figure by Hoffmann from material Dillenius had sent to Jacquin, Jørgensen, James and Jarvis argued that the morphology and habitat instead match young, opening apothecia of what is now Parmelia omphalodes. ^[20] They therefore treated Lichen ampullaceus as a later synonym of Lichen omphalodes and designated Dillenius's figure in Historia muscorum as lectotype, with a British Museum specimen collected by James in 1980 as epitype. ^[21] Index Fungorum follows this treatment, placing the name and its homotypic synonyms in the Parmeliaceae. ^[22]
3. Lichen atro-albus L. was introduced in Species plantarum (1753) as a crustose, saxicolous lichen, but no original material is preserved in the Linnaean Herbarium and the brief protologue could apply to several different alpine crusts. ^[30] Acharius applied the name to a member of the Rhizocarpon badioatrum complex, and later authors used it inconsistently for various lecidioid crusts, so its identity remained unclear and the name fell out of use at species rank. ^[30] Jørgensen, James and Jarvis therefore treated L. atro-albus as a species non satis nota without a type and proposed its rejection under Art. 56 of the Code. ^{[15] [21]} The Nomenclature Committee for Fungi recommended this proposal, and Index Fungorum now lists Lichen atroalbus (without the hyphen) as a formally rejected name (nom. rej.). ^{[16] [31]}
4. Lichen atro-virens L. was introduced in Species plantarum (1753) using the same brief diagnosis that Linnaeus had already published in Flora svecica, with no cited synonyms. It was described simply as a saxicolous crust, and no original material survives in the Linnaean Herbarium, so the name has always been difficult to interpret. ^[30] During the nineteenth and early twentieth centuries it was applied to various taxa within the Rhizocarpon geographicum complex, but the diagnosis is so general that it could fit almost any member of that group. ^{[15] [30]} Runemark therefore treated L. atro-virens as a nomen ambiguum, and the name fell out of regular use. ^[15] Jørgensen, James and Jarvis regarded it as a species non satis nota without a type and proposed its rejection under Art. 56 of the Code. ^{[15] [21]} The Nomenclature Committee for Fungi recommended this proposal, and Index Fungorum now lists Lichen atrovirens (without the hyphen) as a formally rejected name (nom. rej.), with the material historically covered by the name placed within the Rhizocarpon geographicum complex. ^{[16] [32]}
5. Lichen cornucopioides L. was published in Species plantarum (1753) using a diagnostic phrase already employed in Flora svecica, with citations to Flora lapponica and several other works. ^[53] Only one sheet in the Linnaean Herbarium (LINN 1273.217) bears the relevant cross-references; it consists of a richly branched thallus of Cladonia squamosa glued over a single narrow-cupped podetium that Jørgensen, James and Jarvis interpreted as a young member of the Cladonia cornuta complex, best referred to subsp. groenlandica. ^[53] Linnaeus, however, described the apothecia of L. cornucopioides as red (coccineus), whereas both this element and the taxa depicted in the cited figures (Cladonia cervicornis subsp. verticillata) have brown apothecia and were probably treated by him as an unnamed variety under Lichen pyxidatus. ^[53] Because of these inconsistencies, Vainio had already recommended abandoning the name, which has seen virtually no use since his 1887 monograph. ^[15] Jørgensen, James and Jarvis lectotypified L. cornucopioides on the central podetium of LINN 1273.217 but regarded it as a species non satis nota whose continued availability threatened the well-established name Cladonia groenlandica when treated at species rank, and therefore proposed its rejection under Art. 56 of the Code. ^{[15] [53]} The Nomenclature Committee for Fungi supported this proposal, and Index Fungorum now lists Lichen cornucopioides as a rejected name (nom. rej.) with LINN 1273.217 (central specimen) as lectotype. ^{[16] [54]}
6. Lichen fahlunensis L. was published in Species plantarum (1753) using the same diagnostic phrase that Linnaeus had already employed in Flora svecica, with an added reference to Dillenius's Historia muscorum. Several sheets in the Linnaean Herbarium bear the name "fahlunensis", but only LINN 1273.70 carries numerical annotations in Linnaeus's hand that link it to the accounts of L. fahlunensis in Flora svecica and Species plantarum, and its material matches the original description; Jørgensen, James and Jarvis therefore selected it as lectotype. ^[60] The lectotype belongs to the taxon now known as Melanelia stygia (≡ Parmelia stygia (L.) Ach.), and Schaerer later adopted the epithet "fahlunensis" when uniting L. fahlunensis with L. stygius, a choice that would normally have to be followed under Art. 11.5 of the Code. ^{[15] [60]} However, "fahlunensis" has long been used instead for the species now called Cetraria commixta (and in older literature also for C. hepatizon), so reintroducing it for Melanelia stygia would cause serious nomenclatural confusion. ^{[15] [60]} Jørgensen, James and Jarvis therefore proposed Lichen fahlunensis for rejection under Art. 56; the Nomenclature Committee for Fungi recommended this proposal, and Index Fungorum now treats L. fahlunensis as a rejected name (nom. rej.) with LINN 1273.70 as lectotype. ^{[16] [61]}
7. Lichen jubatus L. was published in Species plantarum (1753) using an unchanged diagnostic phrase from Flora svecica, with a citation to its earlier treatment in Flora lapponica and several additional synonyms. ^[66] Two sheets in the Linnaean Herbarium bear this name; Howe selected LINN 1273.281 as lectotype, and Hawksworth later restricted this typification to the left-hand specimen, which carries Linnaeus's annotations linking it to both works. ^[67] Jørgensen, James and Jarvis interpreted this specimen as a coarse, sparsely branched form of what is now treated as Bryoria cf. fremontii, lacking the characteristic yellow soralia, rather than the taxon understood as Alectoria prolixa by Motyka. ^[68] Because Lichen jubatus had been applied inconsistently to various brown pendent Bryoria species, Hawksworth and Sherwood proposed its rejection; this was recommended by the Committee for Fungi and Lichens, and the name is now treated as formally rejected (nom. rej.) under Art. 56 of the Code. ^[66] Index Fungorum lists Lichen jubatus as a rejected name with LINN 1273.281 (left-hand specimen) as lectotype and places the corresponding taxon in the Parmeliaceae. ^[69]
8. Lichen plicatus L. was published in Species plantarum (1753) with a diagnostic phrase borrowed from Flora svecica, cited again from Materia medica and Flora lapponica with additional synonyms. ^[74] No specimen in the Linnaean Herbarium is explicitly labelled with this name; one sheet annotated by Linnaeus as Lichen hirtus and bearing the Flora svecica number of L. plicatus shows that part of his original concept was later absorbed into L. hirtus, while another sheet interpreted by Vainio and Howe as L. plicatus does not fit Linnaeus's broadened later usage. ^[74] Once the Flora svecica material is reassigned to L. hirtus, the only remaining original element for L. plicatus is Dillenius's illustration in Historia muscorum, which depicts Usnea ceratina, a southern Swedish species rather than a Lapland plant. ^{[15] [74]} Because Linnaeus's description is imprecise, the later combination Usnea plicata (L.) F.H.Wigg. was applied to several different taxa in the difficult genus Usnea, and Motyka attempted to revive it for a very rare and poorly known central Swedish entity. ^{[74] [15]} Jørgensen, James and Jarvis lectotypified L. plicatus on Dillenius's figure, thereby tying the name to Usnea ceratina, but argued that the confused history of usage and the threat this posed to the well-established name U. ceratina made it preferable to reject L. plicatus under Art. 56 of the Code. ^{[15] [75]} Their proposal was recommended by the Nomenclature Committee for Fungi, and Index Fungorum now treats Lichen plicatus as a rejected name (nom. rej.) with the Dillenian plate as lectotype. ^{[16] [76]}
9. Lichen roccella L. was introduced in Species plantarum (1753) for a maritime, fruticose lichen used as a dye. Jørgensen, James and Jarvis located two relevant specimens on LINN 1273.263 and, following Howe's earlier choice of that sheet, restricted the lectotype to the lower specimen, which contains lecanoric acid and represents the sorediate Macaronesian taxon they treated as Roccella patellata (≡ R. tuberculosa var. vincentina). ^[79] Candolle later introduced Roccella tinctoria DC. as a replacement name (nomen novum) for Lichen roccella, explicitly citing Linnaeus's species, so the two names share the same type; Tehler showed that R. tinctoria is the correct current name for this sorediate species, with R. patellata, R. vincentina and several other names falling into its synonymy. ^[80] Some modern databases list Roccella phycopsis as the current name for Lichen roccella, but that combination is based on Lichen fucoides Dicks. and applies to a different Atlantic species; Tehler clarified that R. phycopsis is the correct name for R. fucoides, not for Linnaeus's Lichen roccella. ^{[79] [80] [81]}
10. Lichen subfuscus L. was described in Species plantarum (1753) using a diagnostic phrase borrowed from Flora svecica, with "leprosus" altered to "crustaceus", and a single synonym from Dillenius's Historia muscorum. No original material in the Linnaean Herbarium bears this name. Motyka later designated a neotype in the Acharius herbarium (UPS), but because Dillenius's figure is original material available to Linnaeus, it has priority for lectotypification. ^[86] Brodo and Vitikainen accordingly selected Dillenius's plate XVIII, figure 16 as lectotype and identified the corresponding Dillenian specimen as part of the Lecanora subfusca complex, most plausibly referable to Lecanora horiza, although earlier authors had treated it as L. allophana. ^{[86] [87]} Since its introduction, Lichen subfuscus / Lecanora subfusca has been used in a broad and ambiguous way for both corticolous and saxicolous lichens, giving rise to around eighty infraspecific taxa; workers have disagreed over which element represented the "typical" taxon, and Magnusson and others largely abandoned the name as unworkable. ^[87] Brodo and Vitikainen therefore proposed that L. subfuscus be treated as a long-persistent nomen ambiguum and placed on the list of rejected names, arguing that reinstating it in place of the well-established L. allophana would replace a stable name with a confused one. ^[87] Their proposal was recommended by the Nomenclature Committee and approved by the General Committee, and Index Fungorum now treats Lichen subfuscus as a rejected name (nom. rej.) with the Dillenian figure as lectotype. ^{[88] [89]}
11. Mucor fulvus L. was introduced in Species plantarum (1753) with a new diagnostic phrase and no cited synonyms. No original material is preserved in the Linnaean Herbarium, and the description is too imprecise to identify the species with confidence. ^[93] Linnaeus clearly intended a yellow calicioid lichen, and Zahlbruckner treated the name as a form of what is now Chaenotheca furfuracea, but the epithet "fulvus" suggests a more tawny yellow-brown thallus, such as in Chaenotheca chrysocephala, so its identity remains uncertain. ^{[15] [93]} Jørgensen, James and Jarvis therefore regarded M. fulvus as a species non satis nota not in current use and proposed it for rejection under Art. 56 of the Code. ^{[15] [94]} The Nomenclature Committee for Fungi supported this proposal, and Index Fungorum now lists Mucor fulvus as a rejected name (nom. rej.), treating it as an obsolete synonym of Chaenotheca furfuracea and citing a Solander collection from Sweden as lectotype. ^{[16] [95]}
12. Mucor lichenoides L. was published in Species plantarum (1753) using a diagnostic phrase already employed in Flora svecica, with "lentiformi" altered to "lenticulari" and a single synonym from Dillenius's Historia muscorum. Linnaeus left no annotated material in the Linnaean Herbarium, but the Dillenian specimen corresponding to the cited figure consists of two calicioid lichens, Calicium viride and C. calicioid salicinum. ^[97] Because Linnaeus's protologue does not mention a green thallus, Jørgensen, James and Jarvis interpreted the name via the smaller, lower specimen, which has a grey thallus, matches the illustration and represents C. calicioid salicinum; Dillenius had reversed the thallus colours of the two species in his text. ^[97] They therefore selected Dillenius's plate XIV, figure 3 (lower specimen) as lectotype, making the later combination Calicium lichenoides (L.) Schumach. the oldest available name for this species under the Code. ^{[94] [15]} The epithet lichenoides fell out of use early in the twentieth century, whereas Calicium salicinum Pers. has long been the accepted and well-established name for the species. Reintroducing C. lichenoides in place of C. salicinum would replace a stable name with one largely abandoned, so Jørgensen, James and Jarvis proposed Mucor lichenoides for rejection under Art. 56. ^[15] Their proposal was recommended by the Nomenclature Committee for Fungi and accepted by the General Committee, and Index Fungorum now lists Mucor lichenoides as a rejected name (nom. rej.), with the Dillenian figure as lectotype and Calicium salicinum retained as the current name in use. ^{[16] [98]}
13. Mucor sphaerocephalus L. was described in Species plantarum (1753) using an unchanged diagnostic phrase from Flora svecica and with a citation to Haller's 1717 illustration. ^[97] No original specimens are preserved in the Linnaean Herbarium, and Linnaeus's account in Flora svecica shows that he lumped together several discordant elements, including a fungus on Mesembryanthemum in the Uppsala botanical garden and material from walls and stones. ^[97] Haller's plate depicts two quite different organisms: figure 3A, with a grey, fluffy, spherical head, most likely a myxomycete (or possibly a Coniocybe), and figure 3B, with a cup-shaped blackish head, a Calicium that does not match Linnaeus's description of grey-headed (capitula cineria) plants. ^{[15] [97]} Linnaeus therefore seems to have intended the myxomycete-like figure 3A, but without original material its identity cannot be determined with certainty, and the name never came into general use; Acharius already concluded in 1816 that there was no point in maintaining it. ^{[97] [15]} Jørgensen, James and Jarvis treated M. sphaerocephalus as a species non satis nota and proposed the basionym for rejection under Art. 56, a proposal later recommended by the Nomenclature Committee for Fungi and accepted by the General Committee. ^{[94] [15] [16]} Index Fungorum now lists Mucor sphaerocephalus as a rejected name (nom. rej.) with Haller's figure as lectotype and associates it with the well-established calicioid lichen Calicium lenticulare. ^[100]

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