Lydda (planthopper)

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Lydda
Lydda sp.png
A species of Lydda, probably Lydda elongata
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hemiptera
Suborder: Auchenorrhyncha
Infraorder: Fulgoromorpha
Family: Derbidae
Subfamily: Otiocerinae
Tribe: Zoraidini
Genus: Lydda
Westwood, 1840 [1]
Synonyms [2] [3] [4] [5]

Lydda is a genus of planthoppers from the family Derbidae, tribe Zoraidini, with 21 species, as of 2024. [5] The known distribution range of its species is restricted to parts of Queensland and the Northern Territory in Australia, to the island of New Guinea and to the island of Larat in south-eastern Indonesia. [5] [10] The type species, Lydda elongata (Fabricius 1781), is mainly found in parts of Queensland and in New Guinea. [11] In the past, the genus was often confused with the genus Proutista and a number of Lydda species have been originally described under Proutista. While species of Lydda and Proutista share a similar club-shaped outline of their forewings and a similar wing venation, there are significant differences between both genera in the forewing venation and the structure of the aedeagus in males. On the forewings, the media vein has only 5 branches, while Proutista species have 6 branches. In the type species, Lydda elongata, the eyes are unusually large and the head is very broad. The hind wings are about half as long as the forewings. [5] [10] Some species of Lydda have elaborate patterns on their body and forewings. For example, in live specimens of Lydda elongata, the body has a mixture of reddish brown and bluish patterns, while the forewings are brownish with darker spots. [12] Species that have been previously described under the genus Lydda in Africa are now assigned to the genus Lyddastrombus . [13] [5] [14]

Contents

Type species: Cicada elongata Fabricius 1781, a synonym of Lydda elongata

Distribution

Distribution ranges of the species in the genus Lydda (blue outline) and that of Lydda elongata, the type species (brown outline) Lydda distribution2.png
Distribution ranges of the species in the genus Lydda (blue outline) and that of Lydda elongata, the type species (brown outline)

The species of the genus Lydda live in tropical rainforests of north-eastern Australia (some areas in the states of Queensland and the Northern Territory), in all parts of the island of New Guinea (Papua New Guinea and several Indonesian provinces), as well as on the Indonesian island of Larat south of the island of New Guinea. [15] [5] [10] The type species, Lydda elongata, is found mainly in some areas of Queensland, Australia and in some parts of New Guinea. There are additional small, isolated pockets in the Northern Territory of Australia, north of the city of Darwin. [11] While most species of Lydda have a restricted distribution, a few species are more widely distributed. For example, the species Lydda lumholtzi has been recorded from different parts of the island of New Guinea as well as from some areas in northern Australia (Queensland and Northern Territory). [16] Previous distribution records of Lydda species in Africa refer to species now assigned to the genus Lyddastrombus. [13] [5]

Description

Head and thorax of Lydda elongata. Left: dorsal view of head and thorax, right: frontal view of head. These are the original drawings by Kirkaldy of Philadelpheia pandani, a synonym of Lydda elongata Lydda elongata (Kirkaldy - head).png
Head and thorax of Lydda elongata. Left: dorsal view of head and thorax, right: frontal view of head. These are the original drawings by Kirkaldy of Philadelpheia pandani, a synonym of Lydda elongata

The genus Lydda is similar in appearance to the genus Proutista . Both genera have been frequently confused in the past and various species of Lydda have been originally described as species of Proutista. Lydda species can be recognized by a combination of different characters of which the forewing venation and the structure of the aedeagus of males is regarded as most significant. [5] In Lydda elongata, the type species, the head with the eyes is almost as broad as the pronotum in dorsal view, a character which was previously considered to be significant. [17] However, in other species, the head is less broad. [5] The face (frons) of Lydda species is narrow with the ridges on its sides running approximately parallel to each other. In side view, the head has a flat profile. The antennae are short. On the hind legs, the tibia has small lateral spines. The body from the tip of the head to the end of the abdomen is 3–4 mm long. [5]

Shape and venation of forewing (top) and hind wing (bottom) of Lydda elongata, the type species of the genus Lydda Lydda elongata wings.png
Shape and venation of forewing (top) and hind wing (bottom) of Lydda elongata, the type species of the genus Lydda

The forewings are between 6½ and 8½ mm long, while the hind wings are slightly less than half as long as the forewings. The shape of the forewings resembles that of the species of Proutista. Their outline is often club-shaped with the basal third being narrow and the distal part being unusually broad. The venation of the forewings is also similar to that in the genus Proutista with none of the media branches forking further into sub branches. However, among other differences, there are only 5 branches of the media vein in Lydda compared to 6 in Proutista. [5] Similar to the genus Proutista, live specimens of the genus Lydda raise their wings above the body and spread them out at an angle of about 60°. [5] [12] [18]

Lydda species can have elaborate patterns on head and thorax and can be rather colourful. In live specimens of Lydda elongata, the type species, the dorsal parts of head and thorax are partly reddish brown and partly bluish (see image in the taxobox). The face, the facial ridges and the antennae are light blue, as is the central part of the pronotum. These bluish parts are lined with broad, reddish brown stripes and the sides of the head and the pronotum are also partly bluish and partly reddish brown. The mesonotum is reddish brown with bluish longitudinal stripes and the forewings are brownish with darker spots. [12] In Lydda lumholtzi, the forewings are checkered. The thorax is mainly brown with a whitish, trident-shaped mark on the mesonotum. [16] [18]

Biology

Group of Lydda elongata hoppers feeding on a leaf Lydda elongata iNturalist.jpg
Group of Lydda elongata hoppers feeding on a leaf

There is no information on the biology of the nymphal stages of Lydda species. However, it can be assumed that they live in decaying organic matter like other derbids, feeding on fungi. [15] [19] Like for many other derbids, the host plants of the adult hoppers are mainly monocots and include bananas, sugar cane, maize and palm trees. Lydda gressitti and Lydda lurida have been recorded from various host plants in New Guinea, and Lydda elongata has been reported on screw palms (Pandanus) and sugar cane. [16] [5] [11] While the internet contains images of 10 or more insects from the genus Lydda feeding on a leaf, Lydda species are not regarded as harmful to their host plants and most species are rare with a very restricted geographic distribution range. [5] [10]

Taxonomy and species

Westwood described the genus Lydda in 1840 with the type species Derbe elongata from New Holland (Australia), a species originally described by Fabricius in 1781 as Cicada elongata. [1] However, a year later Westwood renamed Lydda as Diospolis. [6] Although the name Diospolis was subsequently used by various authors, the name Lydda is now accepted, based on the priority of the older name. [20] [5] [10] In his descriptions, Westwood did not explain the origin of the names Lydda and Diospolis and why he renamed Lydda as Diospolis. There could be a relationship between these names and the city Lod in Israel, since "Lydda" is the ancient Greek name for that city and "Diospolis" is its name used during the Roman Empire ("Lod" is the Hebrew name). [21]

In 1906, Kirkaldy described the genus Philadelpheia with the type species Philadelpheia pandani from Cairns, Queensland in Australia. [7] However, in 1918, Muir regarded Philadelpheia pandani to be a synonym of Diospolis elongata and therefore Philadelpheia to be a synonym for Diospolis. [4] Philadelpheia is now accepted as a synonym of Lydda and Philadelpheia pandani as a synonym of Lydda elongata (Fabricius, 1781). [20] [5] [10]

In 1907, Distant described the genus Arfaka with the type species Arfaka decida from New Guinea. [8] In 1909, Kirkaldy renamed Arfaka to Afakia, assuming Distant's name was a misspelling for the village "Afaka" in New Guinea. [9] In 1910, Distant clarified that Arfaka was named after the "Arfak" mountains in New Guinea, not after a village. However, he accepted Kirkaldy's new name because the name Arfaka was preoccupied by a genus of Cicadidae. [3] In 1918, Muir regarded both genera Arfaka and Afakia as synonyms of Proutista, but since 1992 they are regarded as synonyms of Lydda. [4] [5] [10]

During a taxonomic revision and a re-description of the genus Lydda in 1992, seven species, described and assigned to Lydda in Africa, were transferred to a new taxon, Diostrombus subgenus Lyddastrombus, with the type species Diospolis annetti Muir, 1918 (= Lydda annetti). [5] Lyddastrombus is now regarded as a full genus, separate from Diostrombus. [14] The removal of the African species from the genus restricts the distribution range of the genus to New Guinea and northern Australia. [5]

The genus Lydda has 21 recognized species as of 2024. [5] None of them seems to be particular common or widespread. The following 6 species have been observed or listed slightly more often, as per Google Scholar and other sources:

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The family Derbidae is a large and diverse group of planthoppers, widely distributed in the tropics and also found in subtropical and temperate regions. It includes more than 150 genera and more than 1500 species. The adults suck the sap of plants while the nymphs live mainly in decaying organic matter, for example under the bark of dead tree trunks, feeding on fungi. In some groups of species the general structure of the insects is largely uniform, whereas in others like in the subfamily Otiocerinae various forms can be found. For example, the shape of the head, the antennae or the wings can differ considerably among genera and species.

<span class="mw-page-title-main">Otiocerinae</span> Subfamily of true bugs

Otiocerinae is a subfamily of planthoppers in the family Derbidae with more than 100 genera and more than 1000 species. The species are widely distributed, especially in the Americas, Sub-Saharan Africa, as well as parts of Asia, Australia and the Pacific. Like in other Derbidae, the adults suck the sap of plants while the nymphs live mainly in decaying organic matter, for example under the bark of dead tree trunks, feeding on fungi.

<span class="mw-page-title-main">Cenchreini</span> Tribe of true bugs

Cenchreini is a tribe of derbid planthoppers in the family Derbidae. There are at least 30 genera in Cenchreini.

<span class="mw-page-title-main">Derbinae</span> Subfamily of true bugs

Derbinae is a subfamily of derbid planthoppers in the family Derbidae.

<i>Proutista moesta</i> Species of planthopper

Proutista moesta is a common species of planthopper from the family Derbidae, tribe Zoraidini. It is found in tropical parts of Asia from India to southern parts of China, some islands in the western Pacific, as well as in the Indian Ocean. There is also a record from eastern Africa, a possible recent introduction. P. moesta can be common on crops like oil palms, coconut palms, sugarcane and several cereals, where it sucks the sap from the plants. It is often regarded as a pest of these crops, a vector of phytoplasma diseases and a biosecurity risk in countries outside its distribution range. Males of P. moesta are smaller than females when measured from the tip of the head to the end of the forewings. The forewings alone have a length of around 6.1 mm for males or around 6.6 mm for females. The colour is predominantly black, but the legs, antennae, facial ridges and the rostrum are yellowish to light orange. The ridges on the mesonotum and the clypeus are whitish to light yellow and the body has scattered white dots. The black forewings have some clear patches, mainly along the costal margin and in the distal half between the hind margin and the media vein. When at rest, the insect raises its wings above the body and spreads them out at an angle of about 60°.

<span class="mw-page-title-main">Rhotanini</span> Tribe of true bugs

The Rhotanini is a group (tribe) of planthoppers in the family Derbidae. They are found mainly in countries of south-eastern Asia like Malaysia, the Philippines, Indonesia and Papua New Guinea. They are characterized by their small size, their broad forewings, the vein pattern on their wings, and a simple head structure. Around 300 species have been described. Like in other derbids, the adults suck the sap of plants while the nymphs live mainly in decaying organic matter.

<i>Levu</i> Genus of planthoppers

Levu is a genus of small planthoppers from the family Derbidae, tribe Rhotanini, with more than 30 species. The species are found mainly on the islands of the western Pacific, particularly Melanesia. A few species have been also reported from eastern Australia and mainland Asia. The adult hoppers are around 4–5 mm long from head to the tip of the forewings. They lack the bright colours or conspicuous wing patters of other derbids, but a few species have red or orange markings. The wings may be carried in a roof-like (tectiform) position when the insects are at rest or they form a common plane like in house flies.

<i>Rhotana</i> Genus of planthoppers

Rhotana is a large genus of planthoppers from the family Derbidae, tribe Rhotanini, with more than 70 species. Most species have been reported from the rainforests of Borneo, Indonesia and New Guinea, but the distribution of the genus extends over a much wider area in southern and south-eastern Asia, as far north as southern Japan and including parts of the Pacific islands as well as Australia. The species are the largest in the tribe Rhotanini, typically between 5–6 mm from head to the tip of the wings. The forewings frequently have striking patterns and colours, some species have also marks on the hind wings.

<i>Saccharodite</i> Genus of planthoppers

Saccharodite is a large genus of planthoppers from the family Derbidae, tribe Rhotanini, with more than 100 species. The largest number of species have been reported from Borneo, Indonesia, New Guinea and the Philippines, but the genus has a much wider distribution, ranging from Sri Lanka in the West, over southern parts of mainland Asia up to southern Japan in the North. Saccharodite is also found in other parts of the western Pacific like Micronesia, Fiji and New Caledonia, up to eastern parts of Australia, in the South as far as Tasmania. The adult hoppers are around 5 mm long from head to the tip of the forewings. They are characterized by their forewing venation. The forewings are usually glassy and bright red marks are common on the head and body, up to most parts except the wings being coloured bright red. When the insects are at rest, the wing surfaces form a common plane like in house flies.

<i>Dichotropis</i> Genus of planthopper

Dichotropis is a small genus of planthoppers from the family Derbidae, tribe Rhotanini, with 9 species. The species are found on the island of New Guinea, including the Indonesian part and the neighbouring islands of New Britain and New Ireland. The distribution range of several species extends to other countries, as far as West Malaysia, the Philippines, the Solomon Islands and northern Australia. The adult hoppers are around 5–6 mm long from head to the tip of the wings. The species can be recognized by their white and powdered forewings which are carried in a roof-like position when the insects are at rest. The ridges on the sides of the face are separated and do not meet in front of the eyes. The body is typically straw-coloured or light brown. The ridges on the head may be orange or the whole head and body may be orange, but bright red marks are missing.

<i>Rhotanella</i> Genus of planthopper

Rhotanella is a small genus of planthoppers from the family Derbidae, tribe Rhotanini, with 8 species. The species are found in the Philippines, Indonesia, New Guinea, the Solomon Islands and southern China. The adult hoppers are between 4–6 mm long from head to the tip of the wings. They can be recognized by the shape and venation of their forewings which have a broad base and 5 very wide costal cells. Especially the first (basal) costal cell is very broad and has a width of about one quarter to one third of the width of the whole wing. The ridges on the sides of the face meet in front of the eyes. The profile of the head is also unusual for the tribe Rhotanini and has either an approximately rectangular or a somewhat triangular outline. The colour of the body is straw-coloured to light brown. The legs may be partly red or have red stripes. The forewings are brownish, but may have red marks in the costal cells. The forewings are carried in a roof-like position when the insects are at rest.

<i>Sumangala</i> (planthopper) Genus of planthopper

Sumangala is a genus of planthoppers from the family Derbidae, tribe Rhotanini. The 13 species are found in southern Asia from southern India and Sri Lanka in the West over parts of mainland Asia to south-east Asian islands like Taiwan and the Philippines and including parts of Indonesia. The adult hoppers are around 5 mm long from head to the tip of the wings. They can be recognized by their forewing venation where the branch of the cubitus joins the first media branch to form the basal median cell, unlike in other genera of Rhotanini where the basal median cell is formed by a crossvein between the first media branch and the cubitus branch or by the first branch of the media splitting again close to its base and the inner sub branch joining the cubitus branch. In Sumangala, the first media branch does not split and as a consequence only 6 veins of the cubitus-media complex reach the margin at the end of the wing. The forewings of Sumangala are mainly whitish and most of them lack larger black marks.

<i>Muiralevu</i> Genus of planthopper

Muiralevu is a genus of planthoppers from the family Derbidae, tribe Rhotanini, with 19 species. While the species of all other genera in the tribe Rhotanini are only found in southern Asia and the Pacific, the species of Muiralevu also occur in western Africa. The distribution of the type species covers several West African countries, ranging from Sierra Leone to the Central African Republic. Four other, very similar species are also found in western Africa, while 14 other species occur in south-eastern Asia. The species of Muiralevu can be recognized by their forewing venation, especially the 4 costal cells and the venation around the basal median cell. Apart from the basal median cell, there is also a characteristic trapezoid cell at the base of the first media branch. On average, the forewings are more elongated compared to most other species of Rhotanini. The adult hoppers are between 4 and 6 mm long from head to the tip of the wings. The head is evenly rounded and the ridges on the sides of the face meet in front of the eyes. The body is usually straw-coloured to light brown. The forewings are mostly whitish and powdered, but in a few species they have large black marks.

<span class="mw-page-title-main">Zoraidini</span> Tribe of true bugs

The Zoraidini is a group (tribe) of planthoppers in the family Derbidae. They are found in tropical and subtropical parts of Africa, Asia, Australia and in parts of the western Pacific. Almost 300 species in 27 genera are recognized. They are characterized by their long and narrow forewings and the short hind wings, usually not longer than half the length of the forewings and often even shorter. The forewings are typically around 8–10 mm long and the body is usually not more than half that length. In most species, the wings are raised when the insects are at rest, and are often spread out at an angle of about 45–90°. The head is compressed and the face is narrow in frontal view. The antennae are often long and the eyes cover about half the face. Like in other derbids, the adults suck the sap of plants while the nymphs live mainly in decaying organic matter. A few species are regarded as minor agricultural pests.

<i>Zoraida</i> (planthopper) Genus of planthoppers

Zoraida is a large genus of planthoppers from the family Derbidae, tribe Zoraidini, with more than 100 species. These are widely distributed in the Old World tropics and in some subtropical parts of eastern Asia. In Africa, they are found mainly in the humid tropics of West Africa like Sierra Leone, the Ivory Coast, the Central African Republic, Nigeria or the Democratic Republic of Congo. However, Zoraida species have been also described from eastern Africa as far north as South Sudan and as far south as parts of South Africa. In Asia, the largest number of species have been described/reported from Indonesia, the Philippines, Taiwan and Sri Lanka, but species are also known as far south as northern Australia and as far north as the southern tip of far-eastern Russia. Like other species of the tribe Zoraidini, Zoraida species have long and narrow forewings and short hind wings. They can be identifies by the forewing venation and the structure of the head.

<i>Proutista</i> Genus of planthoppers

Proutista is a small genus of planthoppers from the family Derbidae, tribe Zoraidini, with 10 species, as of 2024. The type species, Proutista moesta is widely distributed and often very common, its distribution ranging from Tanzania in Africa, over some Indian Ocean islands and southern parts of western Asia to far eastern Asia and parts of the western Pacific. Seven of the remaining species are also found in tropical parts of Asia and only two additional species have been recorded from Africa, where Proutista fritillaris is the most common one. The species of Proutista are characterized by a combination of different features, mainly the shape and venation of the forewings, the size of the hind wings and the structure of the head and the antennae. On the forewings the media vein has 6 branches, none of them branching further into sub branches. The hind wings are about half as long as the forewings and have a rounded tip. The head has a narrow face (frons) and the antennae are rather short, much shorter than the face.

<i>Diostrombus</i> Genus of planthoppers

Diostrombus is a large genus of planthoppers from the family Derbidae, tribe Zoraidini, with more than 40 species. The large majority of species have been described from Africa and the Near East. However, a few species, including the type species, are found in southern and eastern parts of Asia. In Africa, the distribution range of Diostrombus species extends from the humid tropics of western Africa to many parts of eastern Africa, as far north as parts of Sudan and as far south as South Africa. Three species have been described from Yemen and the Iran. Like other species of the tribe Zoraidini, species of Diostrombus have long and narrow forewings and short hind wings. The main characteristic of Diostrombus is the forewing venation with the media vein having 6 branches which do not split further into sub branches. The hind wings are much shorter than half the forewing length and have a pointed tip. In profile the head is usually flat, but in some African species, the ridges on the sides of the face are extended in front of the antennae. The face is narrow. The antennae are short compared to other genera of the tribe Zoraidini, about half as long as the face.

<i>Pamendanga</i> Genus of planthoppers

Pamendanga is a genus of planthoppers from the family Derbidae, tribe Zoraidini, with 37 species, as of 2024. About two-thirds of the species, including the type species, are found in tropical and some subtropical parts of Asia, from Sri Lanka in the West to parts of Japan in the North-East and including the Philippines, Indonesia and New Guinea. The remaining species have been described from Africa, mainly from the humid tropics of West Africa, ranging from Sierra Leone and the Ivory Coast in the North to the Democratic Republic of Congo in the South. Some species are also known from eastern Africa. Like other species of the tribe Zoraidini, Pamendanga species have long and narrow forewings and short hind wings. They can be recognized by the forewing venation and the shape of the head and the antennae. On the forewing, the media vein has 6 branches and the 3rd branch forks further into 2 sub branches. The head has a flat profile and a narrow face with cylindrical antennae which are elongated but not longer than the face.

<i>Lyddastrombus</i> Genus of planthoppers

Lyddastrombus is a small genus of planthoppers from the family Derbidae, tribe Zoraidini, currently (2024) with 8 African species. The morphology of the species is similar to those in the genus Lydda and most species have been originally described as part of Lydda. Lyddastrombus species can be recognized by the venation of their forewings and other features. The genus also shows similarities to the genera Diostrombus and Proutista, sharing with them the short antennae and a related forewing venation, where the media vein has 6 branches with none of them forking further into sub branches. However, Lyddastrombus species differ from both genera by their much broader head. Lyddastrombus was original described as a subgenus of Diostrombus, but it is now regarded as a separate genus, due to the broader head. The species of Lyddastrombus are found mainly in the tropical rainforests of West Africa from Sierra Leone over Nigeria to the Democratic Republic of the Congo, but have been also reported from parts of eastern Africa. Lyddastrombus hoppers have been frequently reported feeding on coconut and oil palms. In Mozambique, they were found to be carriers of a phytoplasma disease of coconut palms.

References

  1. 1 2 Westwood, John O. (1840). "Observations on the genus Derbe of Fabricius". Proceedings of the Linnean Society of London. 1: 82–85.
  2. ↑ Walker, Francis (1851). List of the specimens of homopterous insects in the collection of the British Museum (PDF). Vol. 2. London, England: British Museum. pp. 261–636 [400].
  3. 1 2 Distant, W.L. (1910). "XXX. Rhynchota Malayana. Part III" (PDF). Records of the Indian Museum. 5: 313–338. doi:10.5962/bhl.part.10505.
  4. 1 2 3 Muir, Frederick (1918). "Notes on the Derbidae in the British Museum collection.- I. Zoraidinae". Entomologist's Monthly Magazine. 54: 173–177.
  5. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 Van Stalle, Jan (1992). "Redefinition and revision of Lydda Westwood, 1840 with taxonomic notes on Diostrombus Uhler, 1896 (Homoptera, Fulgoroidea, Derbidae)". Entomologica Scandinavica. 23 (2): 185–213. doi:10.1163/187631292X00290.
  6. 1 2 Westwood, J.O. (1841). "Observations on the genus Derbe of Fabricius". Transactions of the Linnean Society. 19: 1–18. doi:10.1111/j.1096-3642.1842.tb00069.x.
  7. 1 2 Kirkaldy, G.W. (1906). "Leaf-hoppers and their natural enemies (Pt. IX. Leaf-hoppers–Hemiptera)". Bulletin of the Hawaiian Sugar Planters Association. 1 (9): 271–479.
  8. 1 2 Distant, W.L. (1907). "Rhynchotal notes XLII". Annals and Magazine of Natural History. 7. 19: 395–416. doi:10.1080/00222930708562663.
  9. 1 2 Kirkaldy, G.W. (1909). "Hemiptera, old and new, No. 2". The Canadian Entomologist. 41 (11): 388–392. doi:10.4039/Ent41388-11.
  10. 1 2 3 4 5 6 7 "Genus Lydda Westwood, 1840". Australian Faunal Directory. Retrieved 15 October 2024.
  11. 1 2 3 "Species Lydda elongata (Fabricius, 1781)". Australian Faunal Directory. Retrieved 17 October 2024.
  12. 1 2 3 "Lydda elongata". iNaturalist. Retrieved 21 October 2024.
  13. 1 2 Synave, Henri (1973). Monographie des Derbidae africains (Homoptera – Fulgoroidea). Etude du Continent Africain. Vol. 2. Brussels, Belgium: Recherche Fondamentale Collective. pp. 1–223.
  14. 1 2 "Lyddastrombus Van Stalle, 1992". Fulgoromorpha Lists On the Web (FLOW). Retrieved 18 October 2024.
  15. 1 2 Muir, F. (1913). "On some new species of leafhoppers. Part II. Derbidae". Bulletin. Hawaiian Sugar Planters' Association Experiment Station. Division of Entomology. 12: 28–92.
  16. 1 2 3 Van Stalle, Jan (1990). "New species, a key and distribution of the genus Proutista (Homoptera, Derbidae) in New Guinea" (PDF). Indo-Malayan Zoology. 6 (1): 101–118.
  17. ↑ Fennah, R.G. (1952). "On the generic classification of Derbidae (Fulgoroidea), with descriptions of new neotropical species". Transactions of the Royal Entomological Society of London. 103 (4): 109–170. doi:10.1111/j.1365-2311.1952.tb01063.x.
  18. 1 2 "Lydda lumholtzi". iNaturalist. Retrieved 21 October 2024.
  19. ↑ "North American Derbidae". Planthoppers of North America - University of Delaware. Retrieved 4 May 2024.
  20. 1 2 Metcalf, Z.P. (1945). General catalogue of the Hemiptera, Fascile IV, Fulgoroidea. Northampton, Mass., U.S.A.: Smith College. pp. 1–253.
  21. ↑ Dmitriev, Dmitry (2022). "Etymology and grammatical gender of generic names in Auchenorrhyncha (Hemiptera)". Illinois Natural History Survey Bulletin. 43: 1–224. doi: 10.21900/j.inhs.v43.837 .
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