Macrorhizodus Temporal range: | |
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Macrorhizodus praecursor from the Eocene of Kazakhstan | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Chondrichthyes |
Subclass: | Elasmobranchii |
Subdivision: | Selachimorpha |
Order: | Lamniformes |
Family: | Lamnidae |
Genus: | † Macrorhizodus Glikman, 1964 |
Macrorhizodus is an extinct genus of mackerel shark which lived from the early Eocene to early Oligocene epoch of the Paleogene period. It is often considered ancestral to Isurus and sometimes considered part of it. Macrorhizodus is also likely ancestral to Cosmopolitodus . It seems to be related to Isurolamna . It is known from isolated teeth and vertebral centra as well as at least two associated dentitions. [1] It is an incredibly widespread shark, known from every continent except Australia. This includes a report from Antarctica. [2]
The genus is composed of the following species:
Species | Notes | Images | Ref. |
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Macrorhizodus nolfi | This is an early Eocene (Ypresian) species. It is well known from the London Clay of England, the Aktulagay Formation of Kazakhstan, and other places in Eurasia.The species is notably absent in contemporary formations in North America. | [3] | |
Macrorhizodus praecursor | This species is from the mid Eocene to early Oligocene. Some authors restrict this species to the Lutetian stage, while other prefer to lump it with the next two. It is known from the lower Santee Limestone Formation of South Carolina, the Lutetian of Kazakhstan, the Piney Point Formation of Virginia, and other Lutetian localities, as well as from the Rupelian Red Bluff Clay of Alabama. [4] | [1] | |
Macrorhizodus americana | This species is limited to the Bartonian. It is sometimes lumped with M. praecursor. It is best known from Kazakhstan, but has been found in other formations. It is known from the upper Santee Limestone Formation of South Carolina. | [5] [1] | |
Macrorhizodus falcatus | This species is limited to the Priabonian. It is sometimes lumped with M. praecursor. It is common in the Samlat formation of Dakhla, Morocco. | [5] |
This genus appears to have undergone anagenesis, making it difficult to draw lines between species. Some authors prefer to lump and others to split. The early Oligocene species Macrorhizodus flandricus ascribed to this genus now usually considered to be a synonym of Isurus desori, sometimes styled Isurus desori flandrica. However, I. desori might itself be a junior synonym to I. oxyrhincus . This confusion stems from the proximity of Macrorhizodus to Isurus. [1]
The Eocene is a geological epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name Eocene comes from the Ancient Greek Ἠώς and καινός and refers to the "dawn" of modern ('new') fauna that appeared during the epoch.
The Oligocene is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present. As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from Ancient Greek ὀλίγος (olígos) 'few' and καινός (kainós) 'new', and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period.
The Paleogene Period is a geologic period and system that spans 43 million years from the end of the Cretaceous Period 66 million years ago (Mya) to the beginning of the Neogene Period 23.03 Mya. It is the first part of the Cenozoic Era of the present Phanerozoic Eon. The earlier term Tertiary Period was used to define the time now covered by the Paleogene Period and subsequent Neogene Period; despite no longer being recognized as a formal stratigraphic term, "Tertiary" still sometimes remains in informal use. Paleogene is often abbreviated "Pg", although the United States Geological Survey uses the abbreviation "Pe" for the Paleogene on the Survey's geologic maps.
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Carcharodon is a genus of sharks within the family Lamnidae, colloquially called the "white sharks." The only extant member is the great white shark. The extant species was preceded by a number of fossil (extinct) species including C. hubbelli and C. hastalis. The first appearance of the genus may have been as early as the Early Miocene or Late Oligocene.
Megachasma is a genus of mackerel sharks. It is usually considered to be the sole genus in the distinct family Megachasmidae, though suggestion has been made that it may belong in the family Cetorhinidae, of which the basking shark is currently the sole extant member. Megachasma is known from a single living species, Megachasma pelagios.
Paraceratherium is an extinct genus of hornless rhinocerotoids belonging to the family Paraceratheriidae. It is one of the largest terrestrial mammals that has ever existed and lived from the early to late Oligocene epoch. The first fossils were discovered in what is now Pakistan, and remains have been found across Eurasia between China and the Balkans. Paraceratherium means "near the hornless beast", in reference to Aceratherium, the genus in which the type species P. bugtiense was originally placed.
The Priabonian is, in the ICS's geologic timescale, the latest age or the upper stage of the Eocene Epoch or Series. It spans the time between 37.71 and33.9 Ma. The Priabonian is preceded by the Bartonian and is followed by the Rupelian, the lowest stage of the Oligocene.
Isurus is a genus of mackerel sharks in the family Lamnidae, commonly known as the mako sharks. They are largely pelagic, fast predatory fish capable of swimming at speeds up to 50 km/h (31 mph).
Eotheroides is an extinct genus of Eocene sirenian. It is an early member of the family Dugongidae, which includes the extant dugong. Fossils have been found from Egypt, India, and Madagascar. Eotheroides was first described by Richard Owen in 1875 under the name Eotherium, which was replaced by the current name in 1899.
Palaeogale is an extinct genus of carnivorous mammal known from the Late Eocene, Oligocene, and Early Miocene of North America, Europe, and Eastern Asia. A small carnivore often associated with the mustelids, Palaeogale might have been similar to living genets, civets, and linsangs.
Alligator prenasalis is an extinct species of alligator from the Late Eocene period. It is well known, with many fossils having been collected from the Chadron and Brule Formations in South Dakota. The species was first named in 1904, but was originally classified as a crocodile in the genus Crocodilus. It was reassigned to the genus Alligator in 1918 on the basis of more complete material. It is the earliest known member of the genus Alligator.
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Eostegostoma is an extinct genus of prehistoric cartilaginous fish from the Eocene epoch of Belgium and from the Oligocene (Rupelian) of Alabama.
Cosmopolitodus is an extinct genus of mackerel shark that lived between thirty and one million years ago during the late Oligocene to the Early Pleistocene epochs. Its type species is Cosmopolitodus hastalis, the broad-tooth mako. In 2021, Isurus planus was reassigned to the genus, and thus became the second species C. planus. However, some researchers still consider both species of Cosmopolitodus as species of Carcharodon.
Otodus sokolovi is an extinct species or chronospecies of large shark in the family Otodontidae which may represent a transitional chronospecies between Otodus auriculatus and Otodus angustidens. They differ from the former with a less curved root and finer serrations and from the latter with more prominent and recurved cusps. Due to the subtle differences, it is sometimes lumped into O. auriculatus. It, along with the rest of Otodus, is sometimes placed in the genus Carcharocles. Due to its similarities with other chronospecies, it is difficult to tell exactly when it arose and went extinct. Generally, it is said to span from the late Eocene to early Oligocene. They are best known from the late Eocene localities around Dakhla, Morocco and Fayum, Egypt but are represented in many deposits of contemporary age. It measured at least 6 metres (20 ft) long.
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Xiphodolamia is a rare extinct genus of mackerel shark which lived during the Eocene epoch. It is only known from isolated teeth, but has been found in Europe, Africa, and Asia. It is assumed to be pelagic, occurring more frequently in deeper water deposits, most notably the London Clay and Eocene deposits in Denmark. It is distinguished by its rectangular root and twisted blade, unique among mackerel sharks. It is unclear what niche this specialized dentition helped exploit.
Isurolamna is an extinct genus of mackerel shark from the Paleogene period. It contains at least three species and a fourth is sometimes placed in it. It is thought to be closely related to Macrorhizodus, Isurus, and Cosmopolitodus. Isurolamna arose in the Paleocene epoch during the Selandian age, and was extinct by the close of the Rupelian age of the Oligocene epoch. Some features which define this genus include a small, elliptical foramen (hole) in the middle of the lingual side of the root, triangular cusps except on lower interior teeth, and a relatively thick root with nearly vertical margins. The genus has a complicated taxonomic past.