Mesophyletidae Temporal range: | |
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Mekorhamphus beatae paratype | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Infraorder: | Cucujiformia |
Superfamily: | Curculionoidea |
Family: | † Mesophyletidae Poinar, 2008 |
Subfamilies | |
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Synonyms | |
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Mesophyletidae is an extinct family of weevils known from a number of genera preserved in Cretaceous amber. The family was first described as a subfamily in the extant family Caridae, and subsequently raised to family status in 2018.
The family is known exclusively from fossils preserved in Burmese amber, and combined with the Nemonychidae genera, the Burmese amber fauna is considered one of the richest weevil faunas of the Mesozoic, [1] rivaling Kazakhstan's late Jurassic site at Karatau. The Burmese amber fossils preserve more detail than the compression fossils from Karatau. [1] The amber paleoforest was tropical in temperature, and situated along the tidal boundary of a coastal estuary. [2] The resin from which the amber originated is most likely araucarian, based on both inclusions of wood fragments and spectroscopic analysis. [3]
Based on the elongated rostrum, antennae, and ovipositor structures; species in mesophyletidae were likely specialized herbivores which predated seeds and plant ovules in manners similar to that of the living Anthonomini, Curculionini, and related curculionid tribes. The legs are modified with elongated tarsi sporting large claws and tibia with strengthening ridges and a flattened or flared profile. These adaptations indicate an arboreal life of climbing on smooth or flimsy plant organs such as leaves and fruits. [1] Additionally the elytra did not lock along the tips, and are loose to the pygidium which would allow for quick transition to flight, something the group was likely proficient at. They have enlarged eyes composed of coarse facets protruding on their heads, a feature seen in many weevil groups that spend their lives in dark canopies or in leaf litter. [1]
Mesophyletis was the first genus of the group to be described, with its type description being published in 2006. The genus was placed into the family "Eccoptarthridae", an older name sometimes used for Caridae, as one of the genera in the subfamily Mesophyletinae by George Poinar Jr. However, in the 2006 paper, no type genus was specified in the subfamily description and a subsequent paper in 2008 specified Mesophyletis as the type genus. The subfamily Aepyceratinae was proposed in 2017 as a part of the family Nemonychidae based on perceived similarities between Aepyceratus hyperochus and the nemonychids. [1]
The 2018 monograph on Burmese amber weevils expanded the group notably, and raised it to a full family, with two defined subfamilies, Aepyceratinae and Mesophyletinae. An additional five genera were described as part of Aepyceratinae, based on the structuring of the antennae, though the 2018 authors note that there are no other indications that the grouping is monophyletic. [1]
Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small – less than 6 mm in length – and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, with most of them in the family Curculionidae. It also includes bark beetles, which while morphologically dissimilar to other weevils in lacking the distinctive snout, is a subfamily of Curculionidae. Some other beetles, although not closely related, bear the name "weevil", such as the biscuit weevil, which belongs to the family Ptinidae.
The bean weevils or seed beetles are a subfamily (Bruchinae) of beetles, now placed in the family Chrysomelidae, though they have historically been treated as a separate family. They are granivores, and typically infest various kinds of seeds or beans, living most of their lives inside a single seed. The subfamily includes about 1,650 species and are found worldwide.
Belidae is a family of weevils, called belids or primitive weevils because they have straight antennae, unlike the "true weevils" or Curculionidae which have geniculate (elbowed) antennae. They are sometimes known as "cycad weevils", but this properly refers to a few species from the genera Parallocorynus and Rhopalotria.
The family Oedemeridae is a cosmopolitan group of beetles commonly known as false blister beetles, though some recent authors have coined the name pollen-feeding beetles. There are some 100 genera and 1,500 species in the family, mostly associated with rotting wood as larvae, though adults are quite common on flowers. The family was erected by Pierre André Latreille in 1810.
Brentidae, sometimes known as the primitive weevils, is a cosmopolitan family of primarily xylophagous beetles also known as straight-snouted weevils. The concept of this family has been expanded with the inclusion of three groups formerly placed in the Curculionidae; the subfamilies Apioninae, Cyladinae, and Nanophyinae, as well as the Ithycerinae, previously considered a separate family. They are most diverse in the tropics, but occur throughout the temperate regions of the world. They are among the families of weevils that have non-elbowed antennae, and tend to be elongate and flattened, though there are numerous exceptions.
The Bostrichidae are a family of beetles with more than 700 described species. They are commonly called auger beetles, false powderpost beetles, or horned powderpost beetles. The head of most auger beetles cannot be seen from above, as it is downwardly directed and hidden by the thorax. Exceptions are the powderpost beetles, and members of the subfamily Psoinae.
Anthribidae is a family of beetles also known as fungus weevils. The antennae are not elbowed, may occasionally be longer than the body and thread-like, and can be the longest of any members of Curculionoidea. As in the Nemonychidae, the labrum appears as a separate segment to the clypeus, and the maxillary palps are long and projecting.
Nemonychidae is a small family of weevils, placed within the primitive weevil group because they have straight rather than geniculate (elbowed) antennae. They are often called pine flower weevils. As in the Anthribidae, the labrum appears as a separate segment to the clypeus, and the maxillary palps are long and projecting. Nemonychidae have all ventrites free, while Anthribidae have ventrites 1-4 connate or partially fused. Nemonychidae lack lateral carinae on the pronotum, while these are usually present, though may be short, in Anthribidae.
Eobelinae are an extinct weevil subfamily that throve in the late Mesozoic. They belong to the family Nemonychidae. The Eobelidae were widespread, occurring at least in Central Asia, Spain and Brazil during the Late Jurassic and Early Cretaceous.
Astreptolabis is an extinct genus of earwig in the Dermaptera family Pygidicranidae known from a group of Cretaceous fossils found in Myanmar. The genus contains two described species, Astreptolabis ethirosomatia and Astreptolabis laevis and is the sole member of the subfamily Astreptolabidinae.
Tytthodiplatys is an extinct genus of earwig in the family Diplatyidae known from a Cretaceous fossil found in Myanmar. The genus contains a single described species, Tytthodiplatys mecynocercus.
Arostropsis is an extinct genus of broad-nosed weevil in the beetle family Curculionidae known from an Upper Eocene fossil found in Europe. The genus contains a single described species, Arostropsis groehni.
Acanthognathus poinari is an extinct species of ant in the subfamily Myrmicinae known from a single possibly Miocene fossil found on Hispaniola. A. poinari is the first species of the ant genus Acanthognathus to have been described from fossils found in Dominican amber and is one of several species of Acanthognathus found in the Greater Antillas.
Myanmyrma is an extinct genus of ants not placed into any Formicidae subfamily. Fossils of the single known species, Myanmyrma gracilis, are known from the Middle Cretaceous of Asia. The genus is one of several ants described from Middle Cretaceous ambers of Myanmar.
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The amber is of significant palaeontological interest due to the diversity of flora and fauna contained as inclusions, particularly arthropods including insects and arachnids but also birds, lizards, snakes, frogs and fragmentary dinosaur remains. The amber has been known and commercially exploited since the first century AD, and has been known to science since the mid-nineteenth century. Research on the deposit has attracted controversy due to its alleged role in funding internal conflict in Myanmar and hazardous working conditions in the mines where it is collected.
Gerontoformica is an extinct genus of stem-group ants. The genus contains thirteen described species known from Late Cretaceous fossils found in Asia and Europe. The species were described between 2004 and 2016, with a number of the species formerly being placed into the junior synonym genus Sphecomyrmodes.
Enischnomyia is an extinct genus of bat fly in the family Streblidae. At the time of its description the new genus comprised a single species, Enischnomyia stegosoma, known from a single Miocene fossil found on Hispaniola. E. stegosoma was the first fossil streblid bat fly described from a fossil, and the only member of the subfamily Nycterophiliinae described from Hispaniola. The species is host for the plasmodiid Vetufebrus ovatus preserved in its salivary glands and midgut.
2018 in paleoentomology is a list of new fossil insect taxa that were described during the year 2018, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
2017 in paleoentomology is a list of new fossil insect taxa that were described during the year 2017, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.