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Ostracods, or ostracodes, are a class of the Crustacea, sometimes known as seed shrimp. Some 70,000 species have been identified, grouped into several orders. They are small crustaceans, typically around 1 mm (0.039 in) in size, but varying from 0.2 to 30 mm in the case of Gigantocypris. Their bodies are flattened from side to side and protected by a bivalve-like, chitinous or calcareous valve or "shell". The hinge of the two valves is in the upper (dorsal) region of the body. Ostracods are grouped together based on gross morphology. While early work indicated the group may not be monophyletic; and early molecular phylogeny was ambiguous on this front, recent combined analyses of molecular and morphological data found support for monophyly in analyses with broadest taxon sampling.
Traditionally, the Myodocopa and Podocopa have been classified as subclasses within the class Ostracoda, although there is some question about how closely related the two groups actually are. The Myodocopa are defined by possession of a poorly calcified carapace, and 8–9 articles in the exopod of the second antenna. The ventral margin of the carapace is not concave, and the valves do not overlap to a great extent.
Cylindroleberididae is a family of ostracods that shows remarkable morphological diversity. The defining feature is the possession of gills: 7–8 leaf-like pairs at the posterior of the body. Other features common to all species in the family include a "baleen-comb" on both the maxilla and the fifth limb, a sword-shaped coxal endite on the mandible, and the triaenid bristles on the basal endites of the mandible.
The Silurian Lagerstätte preserved in the limestone Wenlock Series of Herefordshire, England, offers paleontologists a rare snapshot of a moment in time, about 420 Mya. In the formation, layers of fine-grained volcanic ash punctuate a sequence of carbonate muds that were accumulating in a marine environment on the outer continental shelf. In this fine-grained matrix, soft-bodied animals and delicate, lightly sclerotized chitinous shells are often preserved in three dimensions, as calcitic fossilizations within calcareous nodules. Calcitic fossilization is an unusual feature.
Megacheira is an extinct class of predatory arthropods that possessed a pair of great appendages, hence the class name as well as the common name "great appendage arthropods". Their neural structures and deutocerebral appendages resemble those of chelicerates. Most of them were found in marine environments throughout the world from the lower to middle Cambrian. Megacheirans were important components of several faunas, including the Burgess, Wheeler and Maotianshan Shales Lagerstatten.
Acaenoplax is an extinct worm-shaped mollusc known from the Coalbrookdale Formation of Herefordshire, England. It lived in the Silurian period. It was a couple of centimetres long and half a centimetre wide, and comprises serially repeated units with seven or eight shells, and rings of 'spines'.
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xiphosura sensu stricto and only regarded as horseshoe crabs under a broader sense. Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata.
Cinerocaris is an extinct genus of crustaceans which existed in the United Kingdom. It contains the species Cinerocaris magnifica.
Bradoriids are an extinct order of small marine arthropods with a bivalved carapace, and were globally distributed, forming a significant portion of the Cambrian and early Ordovician soft-bodied communities.
Pauline avibella is a fossil ostracod from the Silurian with unusually well preserved soft parts, including limbs, eyes, gills and alimentary system.
Pauline is a fossil genus of ostracods from the Silurian. Genus contains two species: Pauline avibella found in 425-million-year-old rocks in the Herefordshire Lagerstätte in England near the Welsh Border and Pauline nivisis, known from the Lower Silurian Pentamerus Bjerge Formation of north Greenland.
Invavita piratica is an extinct, parasitic species of tongue worm, provisionally assigned to the order Cephalobaenida, from Ludlow-aged England. Despite the common name, tongue worms are actually highly modified crustacean arthropods closely related to barnacles and copepods, not worms; the Pentastomida are obligate parasites. It possessed a head, a worm-like body, and two pairs of limbs.
Prosomapoda is a clade of euchelicerates including the groups Xiphosura and Planaterga, as well as several basal synziphosurid genera. The clade is defined by the lack of exopods of prosomal appendage II-V in the adult instar, where in contrast the exopods of appendage II-V are well-developed in the non-prosomapod euchelicerates Offacolus and Dibasterium.
Offacolus is an extinct genus of euchelicerate, a group of chelicerate arthropods. Its only species, O. kingi, has been found in deposits from the Silurian period in the Wenlock Series Lagerstätte of Herefordshire, England. It is the only member of the monotypic family Offacolidae, and classified as a basal ("primitive") genus in the clade Euchelicerata, along with Dibasterium and Prosomapoda. The genus is named after Offa, a king from the ancient kingdom of Mercia, and colus, a person who dwelled among the Offa's Dyke. The species name honors Robert Joseph King, a British mineralogist who found the fossils of Offacolus.
Sollasina is an extinct genus of ophiocistioid that flourished 430 million years ago. Its fossil remains have been recovered from the Coalbrookdale Formation of the Silurian Herefordshire Lagerstätte in England.
Dibasterium is an extinct genus of euchelicerate, a group of chelicerate arthropods. Fossils of the single and type species, D. durgae, have been discovered in deposits of the Middle Silurian period in Herefordshire, England. The name of the genus is derived from the Latin words dibamos and mysterium ("mystery"), meaning "mystery on two legs" and referring to its prosomal limbs. The species name durgae comes from Durga, a Hindu goddess with many arms.
Thanahita is a genus of extinct worm belonging to the group Lobopodia and known from the middle Silurian Herefordshire Lagerstätte at the England–Wales border in UK. It is monotypic and contains one species, Thanahita distos. Discovered in 2018, it is estimated to have lived around 430 million years ago and is the only known extinct lobopodian in Europe, and the first Silurian lobopod known worldwide.
Hallucigeniidae is a family of extinct worms belonging to the group Lobopodia that originated during the Cambrian explosion. It is based on the species Hallucigenia sparsa, the fossil of which was discovered by Charles Doolittle Walcott in 1911 from the Burgess Shale of British Columbia. The name Hallucigenia was created by Simon Conway Morris in 1977, from which the family was erected after discoveries of other hallucigeniid worms from other parts of the world. Classification of these lobopods and their retatives are still controversial, and the family consists of at least four genera.
Coalbrookdale Formation, earlier known as Wenlock Shale or Wenlock Shale Formation and also referred to as Herefordshire Lagerstätte in palaeontology, is a fossil-rich deposit (Konservat-Lagerstätte) in Powys and Herefordshire at the England–Wales border in UK. It belongs to the Wenlock Series of the Silurian Period within the Homerian Age. It is known for its well-preserved fossils of various invertebrate animals many of which are in their three-dimensional structures. Some of the fossils are regarded as earliest evidences and evolutionary origin of some of the major groups of modern animals.
Carbotubulus is a genus of extinct worm belonging to the group Lobopodia and known from the Carboniferous Carbondale Formation of the Mazon Creek area in Illinois, US. A monotypic genus, it contains one species Carbotubulus waloszeki. It was discovered and described by Joachim T. Haug, Georg Mayer, Carolin Haug, and Derek E.G. Briggs in 2012. With an age of about 300 million years, it is the first long-legged lobopodian discovered after the period of Cambrian explosion.
References
- ↑ David J. Siveter; Derek E. G. Briggs; Derek J. Siveter & Mark D. Sutton (2010). "An exceptionally preserved myodocopid ostracod from the Silurian of Herefordshire, UK". Proceedings of the Royal Society B . 277 (1687): 1539–1544. doi:10.1098/rspb.2009.2122. PMC 2871837 . PMID 20106847.
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