Penduline tit

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Penduline tits
Remiz pendulinus 3 (Martin Mecnarowski).jpg
Eurasian penduline tit (Remiz pendulinus)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Infraorder: Passerida
Family: Remizidae
Olphe-Galliard, 1891
Genera

Remiz
Anthoscopus
Auriparus

The penduline tits constitute the family Remizidae, of small passerine birds related to the true tits. All but the verdin make elaborate bag nests hanging from trees (whence "penduline", hanging), usually over water.

Contents

Characteristics

Penduline tits are tiny passerines, ranging from 7.5 to 11 cm in length, that resemble the true tits (Paridae) but have finer bills with more needle-like points. Their wings are short and rounded and their short tails are notched (except the stub-tailed tit). The penduline tits' typical plumage colors are pale grays and yellows and white, though the European penduline tit has black and chestnut markings and some species have bright yellow or red. [1]

Distribution and habitat

The penduline tits live in Eurasia and Africa and North America. The genus Remiz is almost exclusively Palearctic, ranging discontinuously from Portugal and the tip of northern Morocco through to Siberia and Japan. The largest genus, Anthoscopus , is found in sub-Saharan Africa from the Sahel through to South Africa. The verdin lives in arid parts of the southwestern U.S. and northern Mexico.

Several species of penduline tit are migratory, although this behaviour is only shown in species found in Asia and Europe; African species and the verdin are apparently sedentary. The Eurasian penduline tit is migratory over parts of its range, with birds in northern Europe moving south in the winter but birds in southern Europe remaining close to their breeding areas. In contrast the Chinese penduline tit is fully migratory and undertakes long-distance migrations.

Most live in open country with trees or bushes, ranging from desert to marsh to woodland, but the forest penduline tit lives in rain forest. They spend most of the year in small flocks. [1]

Behaviour

Eurasian penduline tit nest in Poland Remiz pendulinis 6.jpg
Eurasian penduline tit nest in Poland

Insects form the larger part of the diet of the penduline tits, and they are active foragers. Their long conical bill is used to probe into cracks and prise open holes in order to obtain prey. Nectar, seeds and fruits may also be taken seasonally. Their foraging behaviour is reminiscent of the true tits (Paridae), foraging upside-down on small branches, manoeuvring branches and leaves with their feet in order to inspect them, and clasping large prey items with one foot while dismembering them. [2]

The common name of the family reflects the tendency of most species to construct elaborate pear-shaped nests. These nests are woven from spiderweb, wool and animal hair and soft plant materials, which are suspended from twigs and branches in trees. The nests of the African genus Anthoscopus are even more elaborate than the Eurasian Remiz, incorporating a false entrance above the true entrance which leads to a false chamber. The true nesting chamber is accessed by the parent opening a hidden flap, entering and then closing the flap shut again, the two sides sealing with sticky spider webs. [2]

Nest of Eurasian penduline tit (Remiz pendulinus) (University of Hamburg) Remiz pendulinus Naumann.png
Nest of Eurasian penduline tit (Remiz pendulinus) (University of Hamburg)

The verdin builds a domed nest out of thorny twigs. In some penduline tit species the eggs are white, sometimes with red spots. The verdin lays blue-green eggs with red spots. Incubation lasts about 13 or 14 days, and the nestlings fledge at about 18 days. [1] In penduline tits, higher incidences[ spelling? ] of extra-pair paternity results in lower rates of male care, suggesting that extra-pair offspring devalues parental care. [3]

Systematics

The family Remizidae was introduced in 1891 (as Remizeae) by the French ornithologist Léon Olphe-Galliard. [4] [5] Sometimes, these birds are included as subfamily Remizinae in the tit family Paridae. Which taxonomic lineup scientists prefers is primarily a matter of taste; that these families are close relatives is well established by now. If they are considered a separate family, the sultan tit and the yellow-browed tit would possibly need to be excluded from the Paridae. [6] [7] The placement of the tit-hylia within this family is particularly controversial, it having variously been placed with the sunbirds, waxbills, honeyeaters and most recently close to the green hylia. [8] It is placed in the family Cettiidae.

The family contains 11 species in 3 genera: [9]

ImageGenusLiving species
Remiz pendulinus (Marek Szczepanek) zoom.jpg Remiz Jarocki, 1819
Southern Penduline-Tit (Anthoscopus minutus).jpg Anthoscopus Cabanis, 1850
Auriparus flavicepsPCCA20050310-5817B.jpg Auriparus S.F. Baird, 1864

Related Research Articles

<span class="mw-page-title-main">Aegithalidae</span> Family of birds

The bushtits or long-tailed tits are small passerine birds from the family Aegithalidae, containing 13 species in three genera, all but one of which (Psaltriparus) are found in Eurasia. Bushtits are active birds with long tails compared to their size, moving almost constantly while they forage for insects in shrubs and trees. During non-breeding season, birds live in flocks of up to 50 individuals. Several bushtit species display cooperative breeding behavior, also called helpers at the nest.

<span class="mw-page-title-main">Tit (bird)</span> Family of small passerine birds

The tits, chickadees, and titmice constitute the Paridae, a large family of small passerine birds which occur mainly in the Northern Hemisphere and Africa. Most were formerly classified in the genus Parus.

<span class="mw-page-title-main">Verdin</span> Genus of birds

The verdin is a species of penduline tit. It is the only species in the genus Auriparus and the only representative of the old world family Remizidae to be found in North America.

<span class="mw-page-title-main">Passerida</span> Clade of birds

Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri. While more recent research suggests that its sister parvorder, Corvida, is not a monophyletic grouping, the Passerida as a distinct clade are widely accepted.

<span class="mw-page-title-main">Sultan tit</span> Species of bird

The sultan tit is an Asian forest bird with a yellow crest, dark bill, black upperparts plumage and yellow underparts. The sexes are similar. The female has greenish-black upperparts and a yellowish throat. The young bird is duller than the adult and has a shorter crest. It is the only member of the monotypic genus Melanochlora, which is fairly distinct from the Parus tits with the nearest relative being the monotypic Sylviparus.

<span class="mw-page-title-main">Eurasian penduline tit</span> Species of bird

The Eurasian penduline tit or European penduline tit is a passerine bird of the genus Remiz. It is relatively widespread throughout the western Palearctic. It is migratory in the northern part of its range but resident in the southern part.

<span class="mw-page-title-main">Grey penduline tit</span> Species of bird

The grey penduline tit, also known as the African penduline-tit, is a species of bird in the family Remizidae. It is found in Angola, Botswana, Burundi, Republic of the Congo, Democratic Republic of the Congo, Eswatini, Kenya, Malawi, Mozambique, Namibia, Rwanda, South Africa, Tanzania, Uganda, Zambia, and Zimbabwe. Its natural habitats are subtropical or tropical dry forests, dry savanna, and moist savanna. At 8 to 9 cm in length and a weight of 6.5 g (0.23 oz), it is one of the smallest species of bird found in Africa, along with its cousin the Cape penduline tit and the perhaps smaller mouse-colored penduline tit and the tit hylia.

<i>Anthoscopus</i> Genus of birds

Anthoscopus is a genus of birds in the penduline tit family Remizidae. The genus is restricted to Sub-Saharan Africa, where it ranges from the Sahel to South Africa. Unlike many of the Eurasian penduline, these species are not generally migratory, instead remaining close to their breeding sites year round. A wide range of habitats is occupied by the six species, from deserts to woodlands to rainforest.

<span class="mw-page-title-main">Cape penduline tit</span> Species of bird

The Cape penduline tit or southern penduline tit is a species of bird in the family Remizidae. It is found in Angola, Botswana, Namibia, South Africa, and Zimbabwe. Its natural habitats are dry savannah, subtropical or tropical dry shrubland, and Mediterranean-type shrubby vegetation. At eight centimetres (3 in) in length, it is one of the smallest species of bird found in Africa, along with its cousins the grey penduline tit and the mouse-coloured penduline tit.

<span class="mw-page-title-main">Mouse-coloured penduline tit</span> Species of bird

The mouse-coloured penduline tit or mouse-colored penduline tit is a species of bird in the family Remizidae. At 8 cm (3.1 in) in length, it is one of the two shortest birds native to Africa, alongside the tit hylia.

<span class="mw-page-title-main">Yellow penduline tit</span> Species of bird

Yellow penduline tit is a species of bird in the family Remizidae. This small yellow passerine bird is found in semi-arid savanna regions of West Africa.

<span class="mw-page-title-main">Fire-capped tit</span> Species of bird

The fire-capped tit is a small, 10 cm (3.9 in) long, weighing about 7 g (0.25 oz) bird species assigned to the family Paridae, that breeds in the temperate forest bordering the Himalayas to the south, in the Hengduan Shan and Nujiang Shan on the Myanmar-China border, the Micah Shan and Daba Shan on the Northern Sichuan border. It winters down hill and further south. Further to the east, birds tend to be smaller and the plumage becomes gradually darker.

<span class="mw-page-title-main">Chinese penduline tit</span> Species of bird

The Chinese penduline tit is a species of bird in the family Remizidae. The species was first described by Robert Swinhoe in 1870. It is found in China, Japan, North Korea, South Korea, and Russia. There are 10 species in the family Remizidae. They are similar to true tits, but make characteristic penduline nests. They inhabit open fields and wetlands. They have sharp bill tips and the sexes look alike.

<i>Remiz</i> Genus of birds

Remiz is a genus of birds in the family Remizidae, commonly known as the Eurasian pendulines. Like other penduline tits, they are named for their elegant, pendulous nests.

<span class="mw-page-title-main">White-crowned penduline tit</span> Species of bird

The white-crowned penduline tit is a species of bird in the family Remizidae. It is found in Afghanistan, China, India, Kazakhstan, Mongolia, Pakistan, Russia, and Tajikistan.

<span class="mw-page-title-main">Black-headed penduline tit</span> Species of bird

The black-headed penduline tit is a species of bird in the family Remizidae. It is found in Central Asia in reed beds along lakes or rivers. Its distribution is fragmented. It is the least thoroughly-documented bird in the Remiz genus, and has been described as one of the most poorly-known bird in Central Asia.

<span class="mw-page-title-main">Hyliidae</span> Family of birds

Hyliidae is a family of passerine birds which contains just two species, the green hylia and the tit hylia. Physiological similarities and molecular phylogenetic studies strongly support the creation of this family.

<span class="mw-page-title-main">Paroidea</span> Superfamily of birds

The Paroidea are a superfamily of passerine birds.

References

  1. 1 2 3 Perrins, Christopher M. (2003). The Firefly Encyclopedia of Birds: Penduline Tits . Firefly Books. ISBN   978-1-55297-777-4.
  2. 1 2 Madge, Steve (2008). "Family Remizidae (Penduline-tits)". In Josep, del Hoyo; Andrew, Elliott; David, Christie (eds.). Handbook of the Birds of the World. Volume 13, Penduline-tits to Shrikes. Barcelona: Lynx Edicions. pp. 52–75. ISBN   978-84-96553-45-3.
  3. Ball, A. D.; van Dijk, R. E.; Lloyd, P.; Pogány, Á.; Dawson, D. A.; Dorus, S.; Bowie, R. C. K.; Burke, T.; Székely, T. (2017). "Levels of extra-pair paternity are associated with parental care in penduline tits (Remizidae)" (PDF). Ibis. 159 (2): 449–455. doi: 10.1111/ibi.12446 .
  4. Olphe-Galliard, Léon (1891). Contributions a la faune ornithologique de l'Europe occidentale (in French). Vol. 3 (fascicles 22–29). Bordeaux, Berlin. p. 64.{{cite book}}: CS1 maint: location missing publisher (link)
  5. Bock, Walter J. (1994). History and Nomenclature of Avian Family-Group Names. Bulletin of the American Museum of Natural History. Vol. 222. New York: American Museum of Natural History. pp. 153, 251. hdl:2246/830.
  6. Jønsson, Knud A.; Fjeldså, Jon (2006). "Determining biogeographical patterns of dispersal and diversification in oscine passerine birds in Australia, Southeast Asia and Africa". J. Biogeogr. 33 (7): 1155–1165. doi:10.1111/j.1365-2699.2006.01507.x. S2CID   84542347.
  7. Gill, Frank B; Slikas, Beth; Sheldon, Frederick H. (2005). "Phylogeny of titmice (Paridae): II. Species relationships based on sequences of the mitochondrial cytochrome-b gene". Auk . 122: 121–143. doi:10.1642/0004-8038(2005)122[0121:POTPIS]2.0.CO;2. S2CID   86067032.
  8. Sefc, Kristina M.; Payne, Robert B.; Sorenson, Michael D. (2003). "Phylogenetic relationships of African sunbird-like warblers: Moho (Hypergerus atriceps), Green Hylia (Hylia prasina) and Tit-hylia (Pholidornis rushiae)" (PDF). Ostrich: Journal of African Ornithology. 74 (1–2): 8–17. doi:10.2989/00306520309485365. S2CID   86085338. Archived from the original (PDF) on 2015-10-31.
  9. Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Waxwings and allies, tits, penduline tits". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved 10 December 2021.

Further reading