River darter | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Perciformes |
Family: | Percidae |
Genus: | Percina |
Species: | P. shumardi |
Binomial name | |
Percina shumardi (Girard, 1859) | |
Synonyms [3] | |
Hadropterus shumardiGirard, 1859 |
Percina shumardi (river darter) is a benthic species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to North America. It is an inhabitant of small and medium rivers where it occurs in rocky riffles with clear, fast-flowing water. The river darter can reach up to 7.3 cm (2.9 in) in length and has an average lifespan of three years. It can be distinguished from other darters by its unique front and rear spots on the dorsal fin.
The river darter is a species native only to the United States and Canada. Within Canada, the river darter is native to southern regions including the Hudson Bay basin, Ontario, and Manitoba. Within the United States, the river darter is native to a band that starts northward in Canada and extends into Minnesota and North Dakota and continues to the Gulf drainages in Alabama and Texas. [2] The river darter is the most common species of darter in the Mississippi River Channel. [4]
Darters are characterized by two dorsal fins and dorsal scutes, a line of modified scales along the midline of the body. They have two anal spins and a complete lateral line. Darters have a moderately pointed snout and a subterminal mouth. River darters are olive brown with distinct dorsal blotching and a mottled back. The dorsal blotches form 8-15 faint bars along the body that become more discernibly ovular towards the tail. A distinctive bar can be seen below the eye. The dorsal fins are clear with light banding. Nuptial males have a bright, golden band in the posterior region of the first, spiny dorsal fin. [5] The second dorsal fin has up to 16 soft rays. The anal fin has 11 rays. [6] Males are darker and more distinctly marked during spawning season, but do not develop bright colors like many other darters. Tubercles develop on the caudal, pelvic and anal fins, and occasionally also occur on the heads of males during the breeding season. [7]
As its name implies, the river darter can be found in major rivers and at the mouths of adjoining tributaries. River darters prefer to inhabit regions with moderate to swift currents in regions around 1 metre (3 ft 3 in) deep. They inhabit chutes of oxbow rivers and riffles containing sandy, gravely, and rocky substrates. During the evening river darters migrate inshore and are commonly found along sandbars in shallower depths of around .6–.9 metres (2 ft 0 in – 2 ft 11 in). During the day they move away from the shore and are most abundant in 1.2 metres (3 ft 11 in) or more of water. [6]
The river darter relies on frictional contact with the substratum to maintain its position while minimizing its energy usage. The river darter's enlarged pectoral fins help create negative lift and increase the friction between its body and the ground to counteract turbidity. As current speed increases, the river darter, like many lotic fishes, increases its frictional contact with the river bottom by releasing gasses from its swim bladder, which increases its density and keeps it near the river bottom. [8]
The abundance of the River darter in the Mississippi River and tributaries indicates that it is not extremely sensitive to turbidity and water pollution. However, the river darter does require deep swift water habitats, which are quickly disappearing with flood control and river impoundment programs. Darters are at risk to pollution due to their small sizes and benthic breeding and feeding habitats that can be heavily modified by pollution. Somewhere between one half and one third of all darter species are considered to be at risk of extinction. In order to protect the river darter, rivers need to be restored and maintained in their natural state with intact flow patterns. River darter abundance is best surveyed in the summer, when water levels are typically at their lowest. [9]
The river darter is an invertivore that primarily feeds during the day. [7] Adult individuals feed on a wide variety of all aquatic invertebrates such as midge and caddisfly larvae, which are common in streams and rivers. Snails are another fundamental aspect of the diet of mature river darters. Chironomids, trichopteras, and small crustaceans are other key elements of the adult diet of river darters. [4] Juvenile river darters typically feed on much smaller prey. Primarily food sources for juveniles include small zooplankton floating in their freshwater habitats. [7]
The river darter attains much of its adult size during the first year of development, and reaches sexual maturity by the age of one. Males tend to develop faster and become larger than females. [7] The average lifespan of a river darter is between 3–4 years.
The river darter spawns from February to May. River darters are brood hiders; they do not guard their eggs. After they have laid their eggs they do not provide parental care. Females bury their fertilized eggs in gravel depressions in areas of slow current. These gravel depressions are called redds. River darter eggs are able to tolerate temperatures between 13–26 °C (55–79 °F). Some reports indicate upstream migration of river darters during the spring. After spawning the fish may move downstream to reach overwintering habitats. The newly hatched larvae may drift downstream as well, and as they grow the darters shift into faster water habitats of mature darters. [6]
The river darter's scientific name comes from the Greek root word Percina or 'little perch' and the word shumardi which was given in honor of its discoverer, Dr. George C. Shumard (1823-1867), a surgeon with the United States Pacific Railroad Survey. [10] [11]
The snail darter is a small species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in East Tennessee freshwater in the United States and in small portions of northern Alabama and Georgia. First recorded in 1973, the snail darter was listed as endangered under the U.S. Endangered Species Act of 1973 by 1975. The species was at the center of a major environmental law controversy that involved a lawsuit seeking to halt the completion of Tellico Dam, which posed a risk of extinction for the snail darter by blocking its migratory route. The case was eventually appealed to the U.S. Supreme Court, which ruled on it in its 1978 decision Tennessee Valley Authority v. Hill.
The leopard darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to the United States, where it can be found only in the Little River drainage in Oklahoma and Arkansas. Its typical habitat is medium to large streams with rubble and boulder substrate. It feeds on small invertebrates on the riverbed and spawns mainly in March and April. It is threatened by impoundment, habitat loss and runoff from agricultural activities. It has never been a common species and has been listed as a threatened species in the United States since 1978, and the International Union for Conservation of Nature lists it as a "vulnerable species".
The rainbow darter is a small species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to North America where it is found in small, fast-moving streams and small to medium-sized rivers. It grows to 2 to 3 inches in length. The species is very sensitive to pollution and silt, staying in clean, pollution-free water. The rainbow darter is easily identified by three dark spots on the back, and blue and orange in the dorsal and anal fins.
The amber darter is a small, endangered species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to the Conasauga River and Etowah River in Georgia and Tennessee in the United States. It typically inhabits riffle areas over gravel and sand bottoms, hiding in aquatic vegetation when present. It feeds on small invertebrates and probably breeds between late fall and early spring. It is an uncommon fish with a small range and the International Union for Conservation of Nature has classified its conservation status as being "endangered".
The tangerine darter or river slick is a small species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in the eastern United States. It grows to a length of 4.3 to 7.1 inches, males being bright orange-red while females are yellow. It is insectivorous, picking insect larvae off aquatic plants and the riverbed, and sometimes rolling small stones over to expose prey. It breeds in late spring and early summer, typically in shallow sandy or gravelly riffles. Pollution and habitat degradation may be a problem in parts of its range, however it is a fairly common fish with a wide range and the International Union for Conservation of Nature has classified its conservation status as being of "least concern".
The Roanoke logperch is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in the Roanoke and Chowan drainages in Virginia and North Carolina in the United States. It inhabits low and moderate-gradient streams and rivers in warm, clear water in mostly unsilted gravel and rubble in runs, pools, and riffles. It is primarily insectivorous. This fish is a federally listed endangered species.
The Conasauga logperch is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the United States. It is one of 184 species of darters in North America. It has been listed as endangered throughout its range with critical habitat under the U.S. Endangered Species Act since August 5, 1985.
The longnose darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the United States.
The channel darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to North America where it typically occurs in the sandy or gravelly shallows of lakes and in small and medium-sized rivers in riffles over sand, gravel or rock bottoms. It is a small fish ranging from 34 to 72 mm in length, olive brown with darker speckles and sometimes with a dark spot below the eye and dark blotches along the flank. It feeds mostly on insect larvae and other small invertebrates and breeds in small streams. This species is listed as threatened by the Canadian Species at Risk Act (SARA) but overall it has a wide range and numerous sub-populations and the International Union for Conservation of Nature has rated it as a "least concern species".
The dusky darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in, but not confined to, both large and small rivers, and shallow creeks, in the eastern, southern, and southeastern United States, particularly the Mississippi River drainage system.
Etheostoma exile, the Iowa darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is, along with about 17 other species of darters, is native to the lakes and streams of Iowa.
The Tippecanoe darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the eastern United States.
The gilt darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It can be found in a number of states in the Mississippi River drainage of the United States although it has been extirpated from some river systems in which it was at one time present, mostly due to siltation and pollution problems. Males are more colorful than females and can grow to a length of about 9 cm (3.5 in). It is a benthic fish that feeds primarily on small aquatic insect larvae. Males form territories during the breeding season in late spring and early summer. Spawning typically takes place at the upper ends of riffles with sandy and gravelly bottoms interspersed with larger cobbles. Some organisations are endeavouring to conserve populations of the gilt darter and re-introduce it to states where the fish has been extirpated but suitable habitat still exists.
Percina kusha, the bridled darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is one of the 324 fish species found in Tennessee.
Percina maculata, the blackside darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is a widespread inhabitant of streams and rivers in the Mississippi River watershed. Like other darters it prefers rocky riffles and sandy runs, but is tolerant of pools and still water as well. It is one of the 324 fish species found in Tennessee.
The blackbanded darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to the river systems of the southeastern United States where it is found in the states of Tennessee, Georgia, Florida, Mississippi, Alabama, and Louisiana. It lives over sandy or gravelly bottoms in smaller rivers and streams and its color varies depending on the different habitats in which it lives. It feeds on small insect larvae and is itself preyed on by larger fish. It spawns between February and June depending on locality. It is generally a common fish throughout most of its range.
Percina phoxocephala, the slenderhead darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in North America in the central Ohio and Mississippi River basins, to northeastern South Dakota and the Lake Winnebago system in Wisconsin, and as far south as the Red River in eastern Oklahoma and northeast Texas, typically in small to medium size rivers. It is a colorful species, with an average length of 6 to 9 centimeters. Males take on a deeper hue during the breeding season. It feeds on insect larvae and other small invertebrates, and spawns between April and June. It is a common fish with a very wide range and the International Union for Conservation of Nature has classified its conservation status as being of "least concern".
The frecklebelly darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It primarily occurs in the upper Kentucky and Green river systems of eastern and central Kentucky and north-central Tennessee, being found mostly in the Cumberland Plateau and Highland Rim regions. The fish gets its name from gets its name from the scattered dark spots on its relatively pale underside. However, a possible more telling characteristic of the species is the systematical arrangement of the spots at the base of the tail as well as the continuous stripes down either side of the back. Total population size is unknown but figured to be relatively large. The frecklebelly darter is a benthic darter that relies on invertebrates as its principal diet. The frecklebelly darter can be found primarily in creeks and small rivers with a moderate gradient and a pool/riffle type flow, and spends most of its time in midwater areas of the stream. The major threats of the frecklebelly darter include decimation through perturbations such as strip mining as well as stream channelization projects. The darter is thought to have a high resilience with minimum population doubling time less than 15 months; it is also believed to have low vulnerability. Frecklebelly darter females contain about 100-300 mature ova depending on size. Fish become mature by age 1. Spawning in this darter species involves egg burial where the male mounts the female and deposits eggs in deep depressions created by receptive females.
The sickle darter is a recently identified freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is closely related to a well-known darter, Percina macrocephala. Discovered in 2007 in the upper Tennessee River drainage, the sickle darter is around 90.5 mm (3.56 in) in length and is a slender bodied, freshwater and benthopelagic fish that most likely feeds on small crayfish and mayflies, since it shares a large mouth and long snout with its sister species. The known distribution of the sickle darter is the upper drainage of the Tennessee River of Tennessee, North Carolina and Virginia. It is distinguished from all other Percina darters except its sister species P. macrocephala by a dark suborbital bar and a black bar subtending a medial black spot on the base of the caudal fin. It stays in mostly rocky, sandy, or silty substrates in clear creeks or small rivers in the Appalachian Mountains. The sickle darter spawns in late winter in gravel shoals. It also has large scales which make it different from macrocephala. It is known to be extirpated from most of its home habitat mainly because of agricultural practices that cause siltation and turbidity which causes most populations of the sickle darter to be widely scattered. But where it is found, it can be observed with regularity. Taking all this into consideration, the sickle darter does not have a formal conservation status under any federal or state law although its closely related species is considered a species of concern by the TWRA. Future management plans should include finding more locations of the sickle darter and decreasing further habitat destruction in known distributions by stating it as a fish of concern by state law.
Percina roanoka, the Roanoke darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to Virginia and North Carolina in the United States and is present in West Virginia, possibly from introduction. It is found in the Roanoke, Tar, and Neuse river systems, typically in small to medium size rivers. It is a colorful species with an average length of 4.3 centimeters (1.7 in). It feeds on insect larvae and other small invertebrates, and spawns in late May and June. It is a common fish with a very wide range and the International Union for Conservation of Nature has classified its conservation status as being of "least concern".