Phacopina

Last updated

†Phacopina
Temporal range: TremadocianFamennian
Reedops deckeri cephalon lateral.jpg
Reedops deckeri , a trilobite of the suborder Phacopina
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Phacopida
Suborder: Phacopina
Struve, 1959
Superfamilies

The Phacopina [1] comprise a suborder of the trilobite order Phacopida. Species belonging to the Phacopina lived from the Lower Ordovician (Tremadocian) through the end of the Upper Devonian (Famennian). [2] The one unique feature that distinguishes Phacopina from all other trilobites are the very large, separately set lenses without a common cornea of the compound eye.

Contents

Habitat

As far as currently known, all Phacopina species were marine bottom-dwellers. [2]

Origin

The Early Ordovician genus Gyrometopus (superfamily Dalmanitoidea, family Diaphanometopidae) is probably close to the common ancestor of the Phacopina. Gyrometopus is phacopid in appearance, but a rostral plate is present, unlike in other Phacopina. However, the rostral plate does not divide the cephalic doublure into a left and right section, but instead the rostral suture defines a semicircle in the frontal ¾ of the doublure. [3]

Description

Zeliszkella torrubiae, a dalmanitid, exhibiting 10 thorax segments Zeliszkella torrubiae oblique lateral.jpg
Zeliszkella torrubiae, a dalmanitid, exhibiting 10 thorax segments
A schizochroal eye of Phacops rana, showing the large (0.5mm) individually set calcite lenses, that are unique in the animal kingdom Phacops rana crassituberculata eye 3.jpg
A schizochroal eye of Phacops rana, showing the large (0.5mm) individually set calcite lenses, that are unique in the animal kingdom

The eyes (if present) consist of very large (0.07mm in Tricopelta breviceps to 0.5mm in Phacops rana ), separately set lenses without a common cornea (so called schizochroal eyes). However, some Phacopina species lack eyes, such as the species of the genus Ductina .

Ductina vietnamica, a phacopid that lacks eyes Ductina vietnamica 2 frontal.jpg
Ductina vietnamica, a phacopid that lacks eyes

The natural fracture lines (sutures) of the head run along the top edges of the compound eye. From the back of the eye these cut to the side of the head (proparian) and not to the back. In front of the eye, the right and left facial sutures connect in front of the inflated glabella and consequently the free cheeks (or librigenae) are yoked as a single piece. The part of the skeleton that is ‘tucked under’ (the doublure) has no sutures crossing it to form a rostral plate.
The thorax has 11 (rarely 10) segments, the side lobes (or pleurae) are furrowed, and the articulating facets distinct. [4] [5]

Systematics

Phacops rana crassituberculata, example of the Phacopoidea superfamily. Note the almost complete merger of the glabella, and the effaced pygidium Phacops rana crassituberulata dorsal.jpg
Phacops rana crassituberculata, example of the Phacopoidea superfamily. Note the almost complete merger of the glabella, and the effaced pygidium
Odontochile sp., example of the Dalmanitoidea superfamily. Note the distinct terminal spine, axis and furrows in the pygidium. Odontochile sp lateral.jpg
Odontochile sp., example of the Dalmanitoidea superfamily. Note the distinct terminal spine, axis and furrows in the pygidium.
A roled-up Acastoides zguilmensis, example of the Acastoidea superfamily. Note the 3 pairs of glabellar lateral lobes. Acastoides zguilmensis cephalon.jpg
A roled-up Acastoides zguilmensis, example of the Acastoidea superfamily. Note the 3 pairs of glabellar lateral lobes.

The Phacopina contain 3 superfamilies and 7 families:

Superfamily Phacopoidea

Superfamily Dalmanitoidea

Superfamily Acastoidea

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Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

<span class="mw-page-title-main">Phacopida</span> Extinct order of trilobites

Phacopida ("lens-face") is an order of trilobites that lived from the Late Cambrian to the Late Devonian. It is made up of a morphologically diverse assemblage of taxa in three related suborders.

<span class="mw-page-title-main">Phacopidae</span> Extinct family of trilobites

Phacopidae is a family of phacopid trilobites that ranges from the Lower Ordovician to the Upper Devonian, with representatives in all paleocontinents.

<i>Phacops</i> Genus of arthropods (fossil)

Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.

<i>Calymene</i> Extinct genus of trilobites

Calymene Brongniart, 1822, is a genus of trilobites in the order Phacopida, suborder Calymenina, that are found throughout North America, North Africa, and Europe in primarily Silurian outcrops. Calymene is closely related to Flexicalymene, and both genera are frequently found enrolled. Calymene trilobites are small, typically 2 cm in length. The cephalon is the widest part of the animal and the thorax usually has 13 segments.

<i>Ductina</i> Genus of extinct trilobites

Ductina is a genus of extinct, small to average sized, eyeless phacopid trilobite, that lived during the Devonian.

Eophacops is a genus of trilobites from the order Phacopida, family Phacopidae. These trilobites lived during the Middle Silurian in what now are North America and North-West Europe. Species assigned to this genus can be distinguished from Phacops by their small size (2–3 cm). Eophacops has very large eyes compared to the rest of its body.

<i>Asaphus</i> Extinct genus of trilobites

Asaphus is a genus of trilobites that is known from the Lower and Middle Ordovician of northwestern Europe.

<i>Gyrometopus</i> Genus of trilobites

Gyrometopus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the Arenig stage of the Ordovician Period, approximately 479 to 472 million years ago.

<span class="mw-page-title-main">Alokistocaridae</span>

Alokistocaridae is a family of ptychopariid trilobites that lived from the Botomian epoch of the Early Cambrian until the Late Cambrian. Alokistocarids were particle feeders and left small furrows which are occasionally preserved. Their remains are found worldwide. Elrathia kingii, one of the most collected trilobites in the world, is a typical alokistocarid.

<i>Conocoryphe</i> Genus of trilobites

Conocoryphe is a genus of primarily eyeless trilobites belonging to the family Conocoryphidae. They lived during the Middle Cambrian period, about 505 million years ago. These arthropods lived on the sea bottom (epifaunal) and lived off dead particulate organic matter.

<i>Odontochile</i> Genus of trilobites

Odontochile is a genus of trilobites in the order Phacopida, family Dalmanitidae.

<span class="mw-page-title-main">Fallotaspidoidea</span>

The ”Fallotaspidoidea” are a superfamily of trilobites, a group of extinct marine arthropods. It lived during the Lower Cambrian (Atdabanian) and species occurred on all paleocontinents except for the Gondwana heartland. A member of this group, Profallotaspis jakutensis, has long been the earliest known trilobite, but recently the redlichiid Lemdadella has been claimed as occurring even earlier.

<span class="mw-page-title-main">Diaphanometopidae</span> Extinct family of trilobites

Diaphanometopidae is a family of trilobites. Its representatives lived during the Arenig and Llanvirn stages of the Ordovician Period, approximately 479 to 463 million years ago. The Diaphanometopidae are thought to have been an early transitional group between the Ptychopariida ancestors and all other Phacopina. Diaphanometopidae, are found in the Lower and Middle Ordovician of Sweden and Russia. Three species are assigned to this family: Diaphanometopus volborthi that has been found in the Dapingian of Baltica, Gyrometopus lineatus occurring in the Floian of Baltica and Prodalmanitina nikolaevi that is known from the Floian of Kolyma. These species have many ancestral characters compared to other Phacopina, but they do not seem to be each other's nearest relatives, which makes it unlikely this family will be maintained when the phylogeny has been studied in more detail.

<span class="mw-page-title-main">Eodiscina</span>

Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.

<i>Pricyclopyge</i> Extinct genus of trilobites

Pricyclopyge is a genus of trilobites assigned to the family Cyclopygidae that occurs throughout the Ordovician. Pricyclopyge had an extratropical distribution, and there is evidence that it lived in darker parts of the water column. Pricyclopyge has huge eyes, an inverted pear-shaped glabella, six thorax segments, with on the 3rd two small discs. Pricyclopyge is known from what are today China, the Czech Republic, France, and the United Kingdom.

Carolinites is a genus of trilobite, assigned to the Telephinidae family, that occurs during the Lower and Middle Ordovician. Carolinites had a pantropical distribution, and there is evidence that it lived in upper parts of the water column. The free cheeks of Carolinites are largely covered by its huge eyes, except for the attachment of large genal spines that extend downward, backward and lateral and gradually curving further backward. The glabella is slightly bulbous, the occipital ring is well defined, but further transglabellar furrows are lacking. The thorax has 10 segments. The axis of the pygidium is highly vaulted, with a curved spine emerging almost perpendicular to the midline and ending parallel to it and a node on each of the other three segments. Carolinites is known from what are today Australia (Tasmania), Canada (Alberta), China, France, Spitsbergen, and the United States (Utah).

<i>Viaphacops</i>

Viaphacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived during the Middle Devonian, and is known from North and South America, Asia.

<span class="mw-page-title-main">Entomaspididae</span> Extinct family of trilobites

Entomaspididae is a family of harpetid trilobites that ranges from the Upper Cambrian to Lower Ordovician of marine strata in China and the United States.

<i>Entomaspis</i> Extinct genus of trilobites

Entomaspis is an extinct genus of harpetid trilobite from Upper Cambrian to Early Ordovician marine strata of the United States. Species are typified by their proportionally large, vaulted, croissant-shaped or bonnet-shaped cephalons that have the cheeks freed to become elongated, curved librigenial spines, and by their comparatively large, crescent-shaped eyes.

References

  1. Struve, W. Suborder Phacopina, p. O461. In: Moore, R.C. (ed.). Treatise on Invertebrate Paleontology. Part O – Arthropoda (Trilobitomorpha). 1959
  2. 1 2 Moore, R.C. (ed.). Treatise on Invertebrate Paleontology. Part O – Arthropoda (Trilobitomorpha). 1959
  3. Jaanusson, V. Evolutionary processes leading to the trilobite suborder Phacopina. Fossils and Strata 4:209–218, fig. 1–5. 1975
  4. Richter, R. Crustacea (Paläontologie). 1933
  5. Whittington, H. B. et al. Part O, Treatise on Invertebrate Paleontology. Revised, Volume 1 – Trilobita – Introduction, Order Agnostida, Order Redlichiida. 1997