Dalmanites

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Dalmanites
Temporal range: Late OrdovicianMiddle Devonian, 449.5–376.1  Ma [1]
Dalmanites limulurus trilobite silurian (cropped).jpg
Specimen of Dalmanites limulurus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Phacopida
Family: Dalmanitidae
Genus: Dalmanites
Barrande, 1852
Species
  • D. caudatus(Bruennich, 1781) (type) = Trilobus caudatus
  • D. corrugatus(Reed, 1901)
  • D. halli(Weller, 1907)
  • D. limulurus(Green, 1832)
  • D. myops(Koenig, 1825)
  • D. nexilus Salter, 1864
  • D. obtusus(Lindstroem, 1885)
  • D. platycaudatusWeller, 1907
  • D. rutellumCampbell, 1967
  • D. socialisBarrande

Dalmanites is a genus of trilobite in the order Phacopida. They lived from the Late Ordovician to Middle Devonian. [1]

Contents

The trilobites of this genus have slightly convex exoskeletons with an average length of 4–7 cm (1.5–3 in). The cephalon is semicircular or parabolic. The glabella (center portion of the head) is often pear-shaped, and tapers outward toward the front. The glabella also always contains three pairs of obvious glabellar furrows. Also prominent are the large mosaic (schizochroal) eyes.

The thorax is composed of eleven segments, with the relatively large pygidium with a slender axis of 11 to 16 rings and 6 or 7 pleural ribs. The pygidium ends in a striking tail spike.

Taxonomy

Species previously assigned to Dalmanites

Some species formerly included in Dalmanites have now been reassigned to other genera.

Description

Specimen with preserved limbs Dalmanites soft tissue.jpg
Specimen with preserved limbs

Dalmanites is genus of trilobites with an average (about 8 centimetres or 3.1 inches long), moderately vaulted exoskeleton with an inverted egg-shaped outline (about 1.5× longer than wide). Its headshield (or cephalon) is semicircular, with robust (genal) spines extending from the side of the cephalon back to approximately the 8th thorax segment. The frontal margin of the cephalon is semicircular to parabolic, and it may have a simple and short anterior extension. The facial suture lies inside or touches the preglabellar furrow. The frontal lobe of the central raised area of the cephalon (or glabella) is much wider than the other lobes. The frontal lobe is vaulted. The eye is moderate to large, about half as long as the cheek. The "seem" that is visible from the ventral side (or doublure) is wide and flat, and has a deep and wide antennal furrow. The "palate" (or hypostome), also only visible from the ventral side, is subtriangular (about as long as wide) and adorned with three weak denticles at its back rim. To the front the hypostome has weak wings extending sideways. The thorax consists of 11 segments. The tips of the segments are pointed and angle back increasingly from about 30° for the anterior segment to slightly pointing inwards for the posterior segment. The tailshield (or pygidium) is large, subtriangular, and about ⅔-¾× as long as wide. The axis is vaulted and ±35% of the width of the pygidium and consists of 12-15 rings. 9–10 deep and wide pleural furrows have flat or only slightly concave bottoms. The furrows within each pleural rib (or interpleural furrows) are very narrow. The frontal band of each pleural rib is more vaulted and broader than the rear band. The pleural furrows almost reach the margin. The pygidial termination (or mucro) is vaulted and more or less pointed into a spine, which may differ between species. The entire exoskeleton is covered in fine and coarse granules. [8] A specimen of an indeterminate species with preserved soft tissue is known from the Silurian aged Coalbrookdale Formation of England. The antennae are not preserved, though biramous limb pairs are preserved, three are present on the cephalon, 11 on the thorax and at least 3 on the pygidium. The thoracic pairs become smaller posteriorly. The exopods of the limbs bear filamentous structures, which appear to be joined with each other by a membrane. [9]

Related Research Articles

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Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

<span class="mw-page-title-main">Redlichiida</span> Extinct order of trilobites

Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.

<span class="mw-page-title-main">Phacopidae</span> Extinct family of trilobites

Phacopidae is a family of phacopid trilobites that ranges from the Lower Ordovician to the Upper Devonian, with representatives in all paleocontinents.

<i>Phacops</i> Genus of arthropods (fossil)

Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.

<i>Paradoxides</i> Extinct genus of trilobites

Paradoxides is a genus of large to very large trilobite found throughout the world during the Middle Cambrian period. One record-breaking specimen of Paradoxides davidis, described by John William Salter in 1863, is 37 cm (15 in). The cephalon was semicircular with free cheeks ending in long, narrow, recurved spines. Eyes were crescent shaped providing an almost 360° view, but only in the horizontal plane. Its elongate thorax was composed of 19–21 segments and adorned with longish, recurved pleural spines. Its pygidium was comparatively small. Paradoxides is a characteristic Middle-Cambrian trilobite of the 'Atlantic' (Avalonian) fauna. Avalonian rocks were deposited near a small continent called Avalonia in the Paleozoic Iapetus Ocean. Avalonian beds are now in a narrow strip along the East Coast of North America, and in Europe.

<span class="mw-page-title-main">Emuellidae</span> Extinct family of trilobites

Emuellidae are a small family of trilobites, a group of extinct marine arthropods, that lived during the late Lower Cambrian of the East Gondwana supercontinent, in what are today South-Australia and Antarctica.

<i>Huntoniatonia</i> Genus of trilobites

Huntoniatonia is genus of trilobites, an extinct group of marine arthropods of average to large size.

<i>Zeliszkella</i> Genus of trilobites

Zeliszkella is a genus of trilobite in the order Phacopida, with species of average size. Species are known from the Middle and Upper Ordovician and have been found in the Czech Republic, France, Morocco, Portugal and Spain.

<i>Dikelocephalus</i> Genus of trilobites

Dikelocephalus is a genus of very large trilobites of up to 50 cm (20 in) long, that lived during the last 3 million years of the Cambrian (Sunwaptan). Their fossils are commonly found as disarticulated sclerites, in the upper Mississippi Valley and in Canada (Alberta). The exoskeleton is rounded anteriorly, with the thorax and sides of the tailshield slightly tapering to about 23× of the width across the base of the spines at the back of the headshield. At the side corners of the pygidium there may be triangular or hooked spines, pointing backwards, while between the spines the posterior margin is at a 30-75° angle with the lateral margin, gently convex or nearly straight. If pygidial spines are lacking, the margin is gradually rounded. The thorax has 12 segments.

<i>Meteoraspis</i>

Meteoraspis is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae. The various species lived from 501 to 497 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Meteoraspis are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as M. nevensis from Victoria Land, Antarctica.

<i>Paralejurus</i> Genus of trilobites

Paralejurus is a genus of trilobite from the Late Silurian to the Middle Devonian of Africa and Europe.

<i>Morocconites</i>

Morocconites malladoides is an average size trilobite, which lived during the Devonian period, in what is now southern Morocco. This species is assumed to be a close relative of Acastoides. The most conspicuous feature is the very long upcurved frontal medial spine, a bit like an avocet bill. It is the only known species in this genus.

The cephalon is the head section of an arthropod. It is a tagma, i.e., a specialized grouping of arthropod segments. The word cephalon derives from the Greek κεφαλή (kephalē), meaning "head".

<i>Biceratops</i>

Biceratops is an extinct genus of olenelloid redlichiid trilobites, of average size, with the largest specimen 8 centimetres or 3.1 inches long, not including the huge pleural spines of the 3rd segment of the thorax. It lived during the Toyonian stage, in what is today the South-Western United States. Biceratops can easily be distinguished from other members of Biceratopsidae by the absence of genal spines, in combination with effaced features of the raised axial area of the head shield, that is bordering the two horn-like projections that carry the eyes. Biceratops nevadensis is the only known species in this genus.

<i>Odontochile</i> Genus of trilobites

Odontochile is a genus of trilobites in the order Phacopida, family Dalmanitidae.

<span class="mw-page-title-main">Eodiscina</span>

Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.

<i>Psychopyge</i>

Psychopyge is a genus of trilobite, that lived during the upper Emsian and has been found in Germany and Morocco. It is characterized by the swordlike extension from the front of the head.

<i>Tricrepicephalus</i>

Tricrepicephalus is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae with species of average size. Its species lived from 501 to 497 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Tricrepicephalus are widespread in Late Cambrian deposits in North America, but is also known from one location in South America. Tricrepicephalus has an inverted egg-shaped exoskeleton, with three characteristic pits in the fold that parallels the margin of the headshield just in front of the central raised area. The articulating middle part of the body has 12 segments and the tailshield carries two long, tubular, curved pygidial spines that are reminiscent of earwig's pincers that rise backwards from the plain of the body at approximately 30°.

Anabaraspis is a genus of redlichiid trilobite, A. splendens occurs in the uppermost Lower Cambrian and lowest Middle Cambrian of Russia. In Anabaraspis, there is an extended area in front of the glabella which is not differentiated in a border and a preglabellar field. It is a unique character in the family Paradoxididae.

<i>Viaphacops</i>

Viaphacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived during the Middle Devonian, and is known from North and South America, Asia.

References

  1. 1 2 Dalmanites at Fossilworks.org
  2. "Dalmanites lapeyrei Bureau 1889". Global Names Index. Archived from the original on 2014-12-16. Retrieved 2014-12-08.
  3. Carvalho, M.G.P.; Fonseca, V.M.M. (2007). "The Trilobite "Dalmanites" maecurua (Middle Devonian, Amazon Basin, Brazil) and the New Genus Amazonaspis (Synphoriidae)". American Museum Novitates (3591): 1–14. doi:10.1206/0003-0082(2007)3591[1:TTDMCM]2.0.CO;2. ISSN   0003-0082. S2CID   198159893 . Retrieved 2014-01-07.
  4. Edgecombe, G.D. (1993). "Silurian Acastacean trilobites of the Americas" (PDF). Journal of Paleontology. 67 (4): 535–548. Bibcode:1993JPal...67..535E. doi:10.1017/S0022336000024884. S2CID   132918487 . Retrieved 2014-01-09.
  5. Lieberman, B.S.; Kloc, G.J. (1997). "Evolutionary and biogeographical patterns in the Asteropyginae (Trilobita, Devonian) Delo, 1935". Bulletin of the American Museum of Natural History. 232. hdl:2246/1623 . Retrieved 16 December 2013.
  6. Corbacho, Joan (2011). "Trilobites from the Upper Ordovician of Bou Nemrou - El Kaid Errami (Morocco) [Trilobites del Ordovícico Superior de Bou Nemrou - El Kaid Errami (Marruecos)]" (PDF). Batelleria. 16: 16–31. Archived from the original (PDF) on 2014-12-20.
  7. Sandford, Andrew C.; Holloway, David J. (2006). "Early Silurian phacopide trilobites from central Victoria, Australia" (PDF). Memoirs of the Museum of Victoria. 63 (2): 215–255. doi:10.24199/j.mmv.2006.63.17. S2CID   130169682.[ permanent dead link ]
  8. Budil, Petr; Thomas, Alan Trevor; Hörbinger, František (2008). "Exoskeletal architecture, hypostomal morphology and mode of life of Silurian and Lower Devonian dalmanited trilobites" (PDF). Bulletin of Geosciences. 83 (1): 2–10. doi: 10.3140/bull.geosci.2008.01.001 . Retrieved 2014-01-07.
  9. Siveter, Derek J.; Fortey, Richard A.; Briggs, Derek E. G.; Siveter, David J.; Sutton, Mark D. (November 2021). Zhang, Xi-Guang (ed.). "The first Silurian trilobite with three-dimensionally preserved soft parts reveals novel appendage morphology". Papers in Palaeontology. 7 (4): 2245–2253. Bibcode:2021PPal....7.2245S. doi: 10.1002/spp2.1401 . ISSN   2056-2799. S2CID   239718706.