Portalia Temporal range: | |
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Holotype specimen of Portalia (right) preserved with Mackenzia (left) | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | incertae sedis |
Genus: | † Portalia |
Species: | †P. mira |
Binomial name | |
†Portalia mira Walcott, 1918 | |
Portalia mira is an extinct species of uncertain placement, described by American paleontologist Charles Walcott from a fossil discovered in the Burgess Shale and initially interpreted as a holothurian echinoderm by Walcott, but this interpretation has been disputed. [1] [2]
The taxonomy of this species remains unresolved. It was initially posited by Walcott that P. mira was a holothurian, but this was disputed by Frizzell and Exline (1955) and Madsen (1957), the latter of whom believed P. mira to be a kind of primitive sponge. J. Wyatt Durham wrote in 1974 that he felt that P. mira should not be rejected as a holothurian without further study. [1] [2] [3]
When examining the fossil holotype, the main body of P. mira is relatively long and contains a central strand that could represent the alimentary canal. Elongated structures resembling tentacles arise along the length of the body and at one end there is an indistinct area that possibly included the head. [1]
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At 508 million years old, it is one of the earliest fossil beds containing soft-part imprints.
Opabinia regalis is an extinct, stem group arthropod found in the Middle Cambrian Burgess Shale Lagerstätte of British Columbia. Opabinia was a soft-bodied animal, measuring up to 7 cm in body length, and had a segmented trunk with flaps along its sides and a fan-shaped tail. The head showed unusual features: five eyes, a mouth under the head and facing backwards, and a clawed proboscis that most likely passed food to its mouth. Opabinia lived on the seafloor, using the proboscis to seek out small, soft food. Fewer than twenty good specimens have been described; 3 specimens of Opabinia are known from the Greater Phyllopod bed, where they constitute less than 0.1% of the community.
Marrella is an extinct genus of marrellomorph arthropod known from the Middle Cambrian of North America and Asia. It is the most common animal represented in the Burgess Shale of British Columbia, Canada, with tens of thousands of specimens collected. Much rarer remains are also known from deposits in China.
Aysheaia is an extinct genus of soft-bodied lobopodian, known from the Middle Cambrian Burgess Shale of British Columbia, Canada
Pikaia gracilens is an extinct, primitive chordate marine animal known from the Middle Cambrian Burgess Shale of British Columbia. Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became "the most famous early chordate fossil", or "famously known as the earliest described Cambrian chordate". It is estimated to have lived during the latter period of the Cambrian explosion. Since its initial discovery, more than a hundred specimens have been recovered.
Anomalocaris is an extinct genus of radiodont, an order of early-diverging stem-group marine arthropods.
Peytoia is a genus of hurdiid radiodont, an early diverging order of stem-group arthropods, that lived in the Cambrian period, containing two species, Peytoia nathorsti from the Miaolingian of Canada and Peytoia infercambriensis from Poland, dating to Cambrian Stage 3. Its two frontal appendages had long bristle-like spines, it had no fan tail, and its short stalked eyes were behind its large head.
Gogia is a genus of primitive eocrinoid blastozoan from the early to middle Cambrian.
Capsospongia, formerly known as Corralia or Corralio, is a middle Cambrian sponge genus known from 3 specimens in the Burgess shale. Its type and only species is Capsospongia undulata. It has a narrow base, and consists of bulging rings which get wider further up the sponge, resulting in a conical shape. Its open top was presumably used to expel water that had passed through the sponge cells and been filtered for nutrients.
Hurdia is an extinct genus of hurdiid radiodont that lived 505 million years ago during the Cambrian Period. Fossils have been found in North America, China, and the Czech Republic.
The fossils of the Burgess Shale, like the Burgess Shale itself, are fossils that formed around 505 million years ago in the mid-Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's 1989 book Wonderful Life describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested.
Eldonia is an extinct soft-bodied cambroernid best known from the Fossil Ridge outcrops of the Burgess Shale, particularly in the 'Great Eldonia layer' in the Walcott Quarry. In addition to over 550 specimens collected by Walcott, 224 specimens of Eldonia are known from the Greater Phyllopod bed, where they comprise 0.43% of the community. Species also occur in the Chengjiang biota, Siberia, and in Upper Ordovician strata of Morocco.
Cambrorhytium is an enigmatic fossil genus known from the Latham Shale (California), and the Chengjiang (China) and Burgess Shale lagerstätte. 350 specimens of Cambrorhytium are known from the Greater Phyllopod bed, where they comprise 0.7% of the community.
Eiffelia is an extinct genus of sponges known from the Middle Cambrian Burgess Shale as well as several Early Cambrian small shelly fossil deposits. It is named after Eiffel Peak, which was itself named after the Eiffel Tower. It was first described in 1920 by Charles Doolittle Walcott. It belongs in the Hexactinellid stem group. 60 specimens of Eiffelia are known from the Greater Phyllopod bed, where they comprise 0.11% of the community.
Mackenzia is an elongated bag-like animal known from the Middle Cambrian Burgess Shale. It attached directly to hard surfaces, such as brachiopod shells. 14 specimens of Mackenzia are known from the Greater Phyllopod bed, where they comprise <0.1% of the community. Mackenzia was originally described by Charles Walcott in 1911 as a holothurian echinoderm. Later, Mackenzia is thought to be a cnidarian and appears most similar to modern sea anemones.
Louisella is a genus of worm known from the Middle Cambrian Burgess Shale. It was originally described by Charles Walcott in 1911 as a holothurian echinoderm, and represents a senior synonym of Miskoia, which was originally described as an annelid. 48 specimens of Louisella are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community. It has been stated to have palaeoscolecid-like sclerites, though this is not in fact the case.
Pagetia is a genus of small trilobite, assigned to the Eodiscinid family Pagetiidae and which had global distribution during the Middle Cambrian. The genus contains 55 currently recognized species, each with limited spatial and temporal ranges.
Redoubtia polypodia is a fossil from the Burgess Shale that was originally described by Charles Walcott as a holothurian echinoderm. Its affinity is unclear, though; it has also been compared to lobopodians.
Mackenziidae is a family of enigmatic animals known from the Cambrian period. Their affinity has been disputed, with the type genus Mackenzia being first described as a holothurian echinoderm, before being moved to a relative of sea anemones, and finally in 2022 moved to Eumetazoa incertae sedis. The family may be related to various members of the Ediacaran biota, due to a similar arrangement of tubular modules.
This is a list of the biota of the Burgess Shale, a Cambrian lagerstätte located in Yoho National Park in Canada.