Pterodactylidae

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Pterodactylids
Temporal range:
Late Jurassic - Early Cretaceous, 152–105  Ma
Pterodactylus antiquus - IMG 0681.jpg
Cast of the sub-adult type specimen of Pterodactylus antiquus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Infraorder: Archaeopterodactyloidea
Clade: Pterodactylidae
Bonaparte, 1838
Subgroups

Pterodactylidae is a controversial group of pterosaurs. During the 2000s and 2010s, several competing definitions for the various Jurassic pterodactyloid groups were proposed. Pereda-Suberbiola et al. (2012) used Fabien Knoll's (2000) definition of the name Pterodactylidae. [1] Knoll had defined Pterodactylidae as a clade containing " Pterodactylus antiquus , Ctenochasma elegans , their most recent common ancestor and all its descendants". [2] Using this definition with the analysis conducted by Pereda-Suberbiola et al. (2012) meant that Ctenochasmatoidea was nested inside Pterodactylidae. [1]

Classification

Below is the majority-rule consensus tree found by Pereda-Suberbiola et al. (2012), showing their preferred definitions of Pterodactylidae and Ctenochasmatoidea. [1]

Pterodactylidae

Pterodactylus

Prejanopterus

Ctenochasmatoidea

Cycnorhamphus

Cearadactylus

Gnathosaurus

Pterodactylus longicolum

Pterodaustro

Ctenochasma

Gegepterus

Eosipterus

Other researchers, such as David Unwin, have traditionally defined Pterodactylidae in such a way to ensure it is nested within Ctenochasmatoidea instead. In 2003, Unwin defined the same clade (Pterodactylus + Pterodaustro) with the name Euctenochasmatia. Unwin considered this to be a subgroup within Ctenochasmatoidea, but most analyses since have found Pterodactylus to be more primitive than he thought, making Euctenochasmatia the more inclusive group. [3]

Related Research Articles

<i>Cearadactylus</i> Genus of anhanguerid pterosaur from the Early Cretaceous

Cearadactylus is a genus of large anhanguerid pterodactyloid pterosaur from the Romualdo Formation of Brazil, South America. Fossil remains of Cearadactylus dated back to the Albian stage of the Early Cretaceous period, about 112 million years ago. The only known species is C. atrox, described and named in 1985 by Giuseppe Leonardi and Guido Borgomanero. The name refers to the Brazilian state Ceará, and combines this with Greek daktylos, "finger", a reference to the wing finger of pterosaurs. The Latin atrox means "frightful", a reference to the fearsome dentition of the species.

<i>Ornithocheirus</i> Genus of ornithocheirid pterosaur from the Early Cretaceous

Ornithocheirus is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the United Kingdom and possibly Morocco.

<span class="mw-page-title-main">Pterodactyloidea</span> Suborder of monofenestratan pterosaurs

Pterodactyloidea is one of the two traditional suborders of pterosaurs, and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal rhamphorhynchoids by their short tails and long wing metacarpals. The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like Nyctosaurus and Tupandactylus. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles.

<span class="mw-page-title-main">Gallodactylidae</span> Family of ctenochasmatoid pterosaurs

Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey.

<i>Lonchodectes</i> Genus of lonchodectid pterosaur from the Late Cretaceous

Lonchodectes was a genus of lonchodectid pterosaur from several formations dating to the Turonian of England, mostly in the area around Kent. The species belonging to it had been assigned to Ornithocheirus until David Unwin's work of the 1990s and 2000s. Several potential species are known; most are based on scrappy remains, and have gone through several other generic assignments. The genus is part of the complex taxonomy issues surrounding Early Cretaceous pterosaurs from Brazil and England, such as Amblydectes, Anhanguera, Coloborhynchus, and Ornithocheirus.

<span class="mw-page-title-main">Lonchodectidae</span> Family of pteranodontoid pterosaurs

Lonchodectidae or Lonchodraconidae is a group of pterosaurs within the clade Pterodactyloidea. It has variously been considered to be within Ctenochasmatoidea, Azhdarchoidea and Pteranodontia. They are notable for their high, conical tooth sockets and raised alveolar margins.

<span class="mw-page-title-main">Ornithocheiroidea</span> Clade of pterodactyloid pterosaurs

Ornithocheiroidea is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods, with fossil remains found all over the world except Antarctica.

<span class="mw-page-title-main">Ctenochasmatoidea</span> Superfamily of archaeopterodactyloid pterosaurs

Ctenochasmatoidea is a group of early pterosaurs within the suborder Pterodactyloidea. Their remains are usually found in what were once coastal or lake environments. They generally had long wings, long necks, and highly specialized teeth.

<span class="mw-page-title-main">Azhdarchoidea</span> Superfamily of ornithocheiroid pterosaurs

Azhdarchoidea is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, Tendaguripterus, that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are Quetzalcoatlus, Hatzegopteryx, and Arambourgiania. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea.

<span class="mw-page-title-main">Dimorphodontidae</span> Family of basal macronychopteran pterosaurs

Dimorphodontidae is a group of early "rhamphorhynchoid" pterosaurs named after Dimorphodon, that lived in the Late Triassic to Early Jurassic. While fossils that can be definitively referred to the group are rare, dimorphodontids may have had a broad distribution, with fossils known from the UK, the southwest United States, and possibly Antarctica.

Prejanopterus is an extinct genus of pterodactyloid pterosaur from the Lower Cretaceous layers of the Leza Formation of La Rioja, Spain.

<span class="mw-page-title-main">Euctenochasmatia</span> Clade of archaeopterodactyloid pterosaurs

Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of Pterodactylus and Ctenochasma, and all their descendants.

<span class="mw-page-title-main">Caelidracones</span> Clade of monofenestratan pterosaurs

The Caelidracones is a group of pterosaurs.

<span class="mw-page-title-main">Germanodactylidae</span> Family of pterodactyloid pterosaurs

Germanodactylidae is a controversial group of pterosaurs within the suborder Pterodactyloidea. It was first named by Yang Zhongjian in 1964, and given a formal phylogenetic definition in 2014 by Brian Andres, James Clark, and Xu Xing. They defined it as the least inclusive clade containing Germanodactylus cristatus and Normannognathus wellnhoferi, which they considered to be close relatives at the time. However, more recent studies by the same researchers have found that these pterosaurs may be only distantly related.

<span class="mw-page-title-main">Eupterodactyloidea</span> Infraorder of pterodactyloid pterosaurs

Eupterodactyloidea is an extinct group of pterodactyloid pterosaurs that existed from the latest Late Jurassic to the latest Late Cretaceous periods. Eupterodactyloids lived on all continents except Antarctica.

<span class="mw-page-title-main">Archaeopterodactyloidea</span> Infraorder of pterodactyloid pterosaurs

Archaeopterodactyloidea is an extinct clade of pterodactyloid pterosaurs that lived from the middle Late Jurassic to the latest Early Cretaceous periods of Africa, Asia, Europe and North America. It was named by Alexander Wilhelm Armin Kellner in 1996 as the group that contains Germanodactylus, Pterodactylus, the Ctenochasmatidae and the Gallodactylidae. In 2003, Kellner defined the clade as a node-based taxon consisting of the last common ancestor of Pterodactylus, Ctenochasma and Gallodactylus and all its descendants. Although phylogenetic analyses that based on David Unwin's 2003 analysis do not recover monophyletic Archaeopterodactyloidea, phylogenetic analyses that based on Kellner's analyses, or the analyses of Brian Andres recover monophyletic Archaeopterodactyloidea at the base of the Pterodactyloidea.

<span class="mw-page-title-main">Novialoidea</span> Clade of macronychopteran pterosaurs

Novialoidea is an extinct clade of macronychopteran pterosaurs that lived from the latest Early Jurassic to the latest Late Cretaceous, their fossils having been found on all continents except Antarctica.

<span class="mw-page-title-main">Ornithocheirae</span> Clade of pteranodontoid pterosaurs

Ornithocheirae is an extinct clade of pteranodontoid pterosaurs from the Early Cretaceous to the Late Cretaceous of Asia, Europe, North America and South America. It was named by Harry Seeley in 1870 as a family that contains Ornithocheirus and its relatives. The name was emended to Ornithocheiridae, to match the requirements of the ICZN Code that a family-ranked clade should end with an -idae suffix. Brian Andres (2010) in his review of pterosaur phylogeny, defined the name Ornithocheirae phylogenetically, as a node-based taxon consisting of the last common ancestor of Anhanguera and Ornithocheirus and all its descendants. Thus Ornithocheirae is defined to include two families, the Anhangueridae and the Ornithocheiridae, following the opinion of Alexander Kellner and Andres that these families should not be synonymized based on their original phylogenetical definitions. However, subsequent studies in 2019 have found Ornithocheirae to be a more inclusive group containing both Anhangueria and Targaryendraconia.

<span class="mw-page-title-main">Tapejaroidea</span> Clade of ornithocheiroid pterosaurs

Tapejaroidea is a group of pterosaurs belonging to the clade Ornithocheiroidea. Tapejaroids lived from the Early to Late Cretaceous periods, with one possible member, Tendaguripterus, extending the fossil range to the Late Jurassic period. Tapejaroidea contains two groups, the Dsungaripteridae and the Azhdarchoidea, which in turn includes the azhdarchids, the group that contains some of the largest flying animals. The group was named by Brazilian paleontologist Alexander Wilhelm Armin Kellner in 1996.

<span class="mw-page-title-main">Ornithocheiromorpha</span> Clade of pteranodontoid pterosaurs

Ornithocheiromorpha is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods, around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing Ornithocheirus, but not Pteranodon.

References

  1. 1 2 3 Xabier Pereda-Suberbiola; Fabien Knoll; José Ignacio Ruiz-Omeñaca; Julio Company; Fidel Torcida Fernández-Baldor (2012). "Reassessment of Prejanopterus curvirostris, a Basal Pterodactyloid Pterosaur from the Early Cretaceous of Spain". Acta Geologica Sinica. 86 (6): 1389–1401. doi:10.1111/1755-6724.12008. hdl: 10651/13364 . S2CID   129917510.
  2. Fabien Knoll (2000). "Pterosaurs from the Lower Cretaceous (?Berriasian) of Anoual, Morocco" (PDF). Annales de Paléontologie. 86 (3): 157–164. doi:10.1016/S0753-3969(00)80006-3.
  3. Unwin, D. M. (2003). "On the phylogeny and evolutionary history of pterosaurs". Geological Society, London, Special Publications. 217 (1): 139–190. Bibcode:2003GSLSP.217..139U. CiteSeerX   10.1.1.924.5957 . doi:10.1144/GSL.SP.2003.217.01.11. S2CID   86710955.