Cycnorhamphus Temporal range: Late Jurassic, | |
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Juvenile specimen | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Ctenochasmatoidea |
Family: | † Gallodactylidae |
Genus: | † Cycnorhamphus Seeley, 1870 |
Species: | †C. suevicus |
Binomial name | |
†Cycnorhamphus suevicus (Quenstedt, 1855) | |
Synonyms | |
Genus synonymy
Species synonymy
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Cycnorhamphus (meaning "swan beak") is a genus of gallodactylid ctenochasmatoid pterosaur from the Late Jurassic period of France and Germany, about 152 million years ago. [1] It is synonymous with the genus Gallodactylus.
In 1855, a fossil in a plate of shale from the Kimmeridgian, found near Nusplingen in Württemberg, holotype GPIT "Orig. Quenstedt 1855, Taf. 1" or GPIT 80, was named Pterodactylus suevicus by Friedrich August Quenstedt. [2] [3] [4] The specific name refers to the tribal area of Suevia. Quenstedt had earlier mentioned the find in a letter to Professor Heinrich Georg Bronn, which was published in 1854. In it he used the name Pterodactylus württembergicus. [5] In 1855 and 1858, Christian Erich Hermann von Meyer adopted this older species name [6] but it would be forgotten afterwards. The publication in 1854 was not meant to be a nomenclatural act. [7] According to Peter Wellnhofer, Pterodactylus württembergicus should be considered a nomen oblitum . [8]
In 1858, Johann Andreas Wagner described a second specimen consisting of the wings, a shinbone and a foot. He named it Pterodactylus (Ornithocephalus) eurychirus, "the broad-handed", but later in the same publication used the name Pterodactylus suevicus eurychirus as if it were a subspecies. [9] This is today considered a junior synonym. The specimen was acquired by the Bayerische Staatssammlung für Paläontologie und Geologie but was lost in April 1944 during the Munich bombardment. [8]
In 1870, Harry Govier Seeley assigned P. suevicus to a new genus: Cycnorhamphus. [10] The name is derived from Greek κύκνος, kyknos, "swan" and ράμφος, ramphos, "beak", in reference to the snout shape. The type species of this genus is Pterodactylus suevicus, the combinatio nova is Cycnorhamphus suevicus.
In 1878, Oscar Fraas referred a specimen of Pterodactylus longicollum , the later Exemplar Nr 58, to Pterodactylus suevicus. [11] Seeley made this the type specimen of yet another species: Cycnorhamphus Fraasii, in 1891. [12]
In 1907 however, Felix Plieninger rejected the split between Pterodactylus and Cycnorhamphus and denied the validity of C. fraasii. [13] This would be the standard interpretation, shared by most paleontologists, for over sixty years.
During the late 1960s, the Ghirardi family began to exploit the chalkstone quarries of Les Besson, located on the French army base of Canjuers near Aiguines. A Lagerstätte proved to be present from which many high quality fossils could be collected. One of these was a slab showing a pterosaur. The precise time and location of this discovery are unknown. It was first reported in the scientific literature in 1971, by Léonard Ginsburg and Guy Mennessier. [14] In 1974, French paleontologist Jacques Fabre based on this specimen, MNHN CNJ-71, named a new species Gallodactylus canjuersensis. [15] The generic name combines a reference to Gallia with Greek daktylos, "finger". He concluded it was the same genus as P. suevicus, but did not revive Cycnorhamphus, judging that the latter name was unavailable because of mistakes in the diagnosis by Seeley, already pointed uit by Plieninger. P. suevicus thus became Gallodactylus suevicus. In 1976, Fabre again named the species, describing it in greater detail but not mentioning the earlier publication. [16] This confused later researchers who mistakenly assumed that 1976 was the formal naming date. In fact the 1974 paper contains a sufficient description and the species was validly named that year. [7] In 1983, the Ghirardis sold their entire collection to the National Museum of Natural History, France.
However, in 1996, Christopher Bennett pointed out that such mistakes do not invalidate a name and that therefore Cycnorhamphus has priority, making Gallodactylus canjuersensisC. canjuersensis. [4] In 2010 and 2012, Bennett published further re-studies of the fossils, concluding that the differences between the two species could be explained by age, sex or individual variation, and formally synonymized C. canjuersensis and C. suevicus. [7]
Cycnorhamphus had historically been assumed to have had long jaws with teeth at the very tip, akin to those of Pterodactylus antiquus . However, recent work on a specimen nicknamed "The Painten Pelican" [7] has revealed that the animal possesses a very unusual jaw anatomy, with peg-like teeth at the jaw tips - blunter and stouter in older individuals -, jaw curvatures behind said teeth that form angled arcs away from the biting surface, forming thus an opening, and two poorly understood soft tissue structures occupying this opening from the upper jaw, showing mineralization. The purpose of these adaptations is unknown, [17] but they are more obvious and well developed in adult animals. It has been speculated that the jaws functioned similar to those of openbill storks, allowing the animal to hold hard invertebrates like mollusks and either crush or bisect them. [18]
As illustrated below, the results of a topology are based on a phylogenetic analysis made by Longrich, Martill, and Andres in 2018. They placed the Cycnorhamphus within the clade Euctenochasmatia, more precisely within the family Gallodactylidae, sister taxon to Normannognathus . [19]
Archaeopterodactyloidea |
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Pterodactylus is an extinct genus of pterosaurs. It is thought to contain only a single species, Pterodactylus antiquus, which was the first pterosaur to be named and identified as a flying reptile and one of the first prehistoric reptiles to ever be discovered.
Rhamphorhynchus is a genus of long-tailed pterosaurs in the Jurassic period. Less specialized than contemporary, short-tailed pterodactyloid pterosaurs such as Pterodactylus, it had a long tail, stiffened with ligaments, which ended in a characteristic soft-tissue tail vane. The mouth of Rhamphorhynchus housed needle-like teeth, which were angled forward, with a curved, sharp, beak-like tip lacking teeth, indicating a diet mainly of fish; indeed, fish and cephalopod remains are frequently found in Rhamphorhynchus abdominal contents, as well as in their coprolites.
Christian Erich Hermann von Meyer, known as Hermann von Meyer, was a German palaeontologist. He was awarded the 1858 Wollaston medal by the Geological Society of London.
Dakosaurus is an extinct genus of crocodylomorph within the family Metriorhynchidae that lived during the Late Jurassic and Early Cretaceous. It was large, with teeth that were serrated and compressed lateromedially. The genus was established by Friedrich August von Quenstedt in 1856 for an isolated tooth named Geosaurus maximus by Theodor Plieninger in 1846. Dakosaurus was a carnivore that spent much, if not all, its life out at sea. The extent of its adaptation to a marine lifestyle means that it is most likely that it mated at sea, but since no eggs or nests have been discovered that have been referred to Dakosaurus, whether it gave birth to live young at sea like dolphins and ichthyosaurs or came ashore like turtles is not known. The name Dakosaurus means "biter lizard", and is derived from the Greek dakos ("biter") and σαῦρος -sauros ("lizard").
Ornithocheirus is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the United Kingdom and possibly Morocco.
Termatosaurus is a genus of archosaur known from several tooth specimens differentiated between two species. Its remains come from the Upper Triassic and was once thought to have survived until the Early Jurassic, but the Jurassic remains were redescribed as plesiosaur remains. It has only been found in France, England, Germany and Switzerland. and two species are known of this animal: the type species, Termatosaurus albertii, named by Meyer and T. Plieninger in 1844; and T. crocodilinus, by Quenstedt (1858). It is very obscure and apparently considered to be dubious.
Germanodactylus is a genus of germanodactylid pterodactyloid pterosaur from Upper Jurassic-age rocks of Germany, including the Solnhofen Limestone. Its specimens were long thought to pertain to Pterodactylus. The head crest of Germanodactylus is a distinctive feature.
Arambourgiania is an extinct genus of azhdarchid pterosaur from the Late Cretaceous period of Jordan, and possibly the United States. Arambourgiania was among the largest members of its family, the Azhdarchidae, and it is also one of the largest flying animals ever known. The incomplete left ulna of the "Sidi Chennane azhdarchid" from Morocco may have also belonged to Arambourgiania.
Ctenochasma is a genus of Late Jurassic ctenochasmatid pterosaur belonging to the suborder Pterodactyloidea. Three species are currently recognized: C. roemeri, C. taqueti, and C. elegans. Their fossilized remains have been found in the Solnhofen Limestone of Bavaria, Germany, the "Purbeck Group" of northeastern Germany, and the Calcaires tâchetés of eastern France.
Campylognathoides is an extinct genus of pterosaur discovered in the Württemberg Lias deposits of Germany; this first specimen however, consisted only of wing fragments. Further better preserved specimens were found in the Holzmaden shale; based on these specimens, Felix Plieninger erected a new genus.
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey.
Ornithostoma is a genus of pterodactyloid pterosaur that lived during the Early Cretaceous period of Europe, around 110 million years ago. Ornithostoma was once thought to have been a senior synonym of the pteranodontid Pteranodon due to its toothless anatomy and prior naming.
Diopecephalus is a genus of pterodactyloid pterosaur from the Lower Tithonian of the Lithographic Limestone, Bavaria, Germany. The type and only species is D. kochi, although the name has been applied to Pterodactylus longicollum, with longicollum erroneously listed as the type species.
Doratorhynchus is a generic replacement name for Pterodactylus validus, in 1875 suggested by Harry Govier Seeley. Today it is considered a nomen vanum.
Altmuehlopterus is a genus of pterosaur belonging to the Pterodactyloidea. It lived in the Late Jurassic of what is now Germany. It was formerly known as "Daitingopterus", a nomen nudum, informally coined in 2004.
Cricosaurus is an extinct genus of marine crocodyliforms of the Late Jurassic. belonging to the family Metriorhynchidae. The genus was established by Johann Andreas Wagner in 1858 for three skulls from the Tithonian of Germany. The name Cricosaurus means "Ring lizard", and is derived from the Greek Krikos- ("ring") and σαῦρος -sauros ("lizard"). It was a relatively small reptile, with C. suevicus and C. araucanensis measuring 2 m (6.6 ft) and 3.2 m (10 ft) in total body length, respectively.
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of Pterodactylus and Ctenochasma, and all their descendants.
Ardeadactylus is an extinct genus of ctenochasmatoid pterosaur known from the Late Jurassic Solnhofen limestone of Bavaria, southern Germany. It contains a single species, Ardeadactylus longicollum, which was originally thought to be a species of Pterodactylus, as P. longicollum.
Aerodactylus is a pterosaur genus containing a single species, Aerodactylus scolopaciceps, previously regarded as a species of Pterodactylus.
Timeline showing the development of the extinct reptilian order Pterosauria from its appearance in the late Triassic period to its demise at the end of the Cretaceous, together with an alphabetical listing of pterosaur species and their geological ages.