Leptostomia Temporal range: | |
---|---|
Life restoration | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Ornithocheiroidea |
Clade: | † Azhdarchoidea |
Genus: | † Leptostomia Smith et al., 2021 |
Type species | |
†Leptostomia begaaensis Smith et al., 2021 |
Leptostomia is a genus of long-beaked pterosaur from the mid-Cretaceous (Cenomanian) of Morocco, North Africa. The type species is L. begaaensis, which was named and described in 2021 from sediments of the Kem Kem Group in Morocco. [1] It was a small animal with a long, slender bill which is thought to have been used to probe sediments for worms and other invertebrates, similar to kiwi birds and curlews. Leptostomia is likely a member of the Azhdarchoidea.
In 2021, paleontologists Roy E. Smith, David Michael Martill, Alexander Kao, Samir Zouhri and Nicholas R. Longrich named the pterosaur species Leptostomia begaaensis. The generic name is a combination of the Ancient Greek leptos (meaning "slim"), and stoma (meaning "mouth"). The specific name refers to the oasis village Hassi El Begaa, near the Aferdou N 'Chaft site where the pterosaur was found. [1]
The holotype specimen of Leptostomia, FSAC-KK 5075, was found in a layer of a river deposit of the Ifezouane Formation of the Kem Kem Group in Morocco. The age of which is uncertain, but it is believed to date back to the Cenomanian or perhaps Albian stages of the middle Cretaceous, about 112-94 million years ago. The holotype consists of part of the upper beak, located in front of the nasoantorbital fenestra. Specimen FSAC-KK 5076, a second fossil of the species, and the paratype specimen of Leptostomia, consists of the anterior part of the dentary symphysis. The preserved portion of the lower jaws may have lain in front of the holotype rostrum according to the authors' interpretation, with some overlap. The specimens of Leptostomia were examined by means of a CT scan. [1]
Leptostomia was a small, long-beaked animal with adaptations for sediment probing. The holotype consists of two beak fragments. The fragments of Leptostomia were mostly flat, and had thick walls, unusual for a pterosaur. The holotype of Leptostomia measured 48 millimeters (1.9 in) in length, and from this measurement, paleontologists have estimated a skull length measuring between 6 and 20 centimeters (2.4 and 7.9 in). [1]
The muzzle of the animal is extremely elongated, only 2.2 millimeters (0.087 in) deep at the front. The jaws are toothless and slightly upcurved, and taper gradually toward the front. The jaws were also strongly flattened. The upper jaw had a narrow ridge, similar to that seen in many other pterosaurs, which would have occluded with a groove in the lower jaw. [1]
Smith et al. suggest that Leptostomia was a pterodactyloid, and likely a member of the clade Azhdarchoidea, but its affinities remain unclear because of the fragmentary nature of the fossils. Because no pterosaurs closely resembling Leptostomia have been described before, it is unclear whether it belongs to a known family such as Azhdarchidae or Chaoyangopteridae, or represents a new azhdarchoid clade. [1]
The beak of Leptostomia is unlike that of any previously described pterosaurs. Instead, it closely resembles birds that feed by probing in sediment, such as kiwis, ibises, hoopoes and snipes. These birds typically feed on invertebrates such as earthworms (in soil) or polychaete worms, fiddler crabs, or bivalves (in estuarine or marine intertidal settings). Leptostomia's prey could have been sought by a sensitive system of nerve endings capable of detecting movement at a distance. [1]
Azhdarchidae is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well. Azhdarchids are mainly known for including some of the largest flying animals discovered, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and Titanopteryx. They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. Previously it was thought that by the end of the Cretaceous, most pterosaur families except for the Azhdarchidae disappeared from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms.
Ornithocheirus is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the United Kingdom and possibly Morocco.
Sigilmassasaurus is a controversial and dubious genus of spinosaurid dinosaur that lived approximately 100 to 94 million years ago during the Late Cretaceous Period in what is now northern Africa. Named in 1996 by Canadian paleontologist Dale Russell, it contains a single species, Sigilmassasaurus brevicollis. The identity of the genus has been debated by scientists, with some considering its fossils to represent material from the closely related species Spinosaurus aegyptiacus, while others have classified it as a separate taxon, forming the clade Spinosaurini with Spinosaurus as its sister taxon.
Thalassodromeus is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of Thalassodromeus sethi by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner", and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to T. sethi in 2005, became the basis of the new genus Banguela in 2015, and assigned back to Thalassodromeus as the species T. oberlii in 2018, though other researchers consider it a valid genus. Another species was described in 2015 based on a supposed crest fragment, but this was later shown to be part of a turtle shell.
Siroccopteryx is an extinct genus of anhanguerid pterodactyloid pterosaur which lived in Morocco during the Cenomanian stage of the Late Cretaceous. Some researchers, such as David M. Unwin, consider the genus a junior synonym of Coloborhynchus.
Coloborhynchus is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. Coloborhynchus is known from the Lower Cretaceous of England, and depending on which species are included, possibly the Albian and Cenomanian ages as well. Coloborhynchus was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related Tropeognathus is now thought to have had a larger wingspan.
Feilongus is an extinct genus of ctenochasmatid pterodactyloid pterosaur from the Barremian–Aptian-age Lower Cretaceous Yixian Formation of Beipiao, Liaoning, China.
Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.
Lonchodectidae or Lonchodraconidae is a group of pterosaurs within the clade Pterodactyloidea. It has variously been considered to be within Ctenochasmatoidea, Azhdarchoidea and Pteranodontoidea. They are notable for their high, conical tooth sockets and raised alveolar margins.
The Cambridge Greensand is a geological unit in England whose strata are earliest Cenomanian in age. It lies above the erosive contact between the Gault Formation and the Chalk Group in the vicinity of Cambridgeshire, and technically forms the lowest member bed of the West Melbury Marly Chalk Formation. It is a remanié deposit, containing reworked fossils of late Albian age, including those of dinosaurs and pterosaurs.
The Kem Kem Group is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations, with the lower Ifezouane Formation and the upper Aoufous Formation used for the strata on the eastern side of the Atlas Mountains (Tinghir), with the Gara Sbaa Formation and Douira Formation used in the southern Tafilalt region. It is exposed on an escarpment along the Algeria–Morocco border.
Chaoyangopteridae is a family of pterosaurs within the larger group Azhdarchoidea. Chaoyangopterids lived mostly during the Early Cretaceous period, though possible members, Microtuban, Xericeps and Argentinadraco, may extend the fossil range to the Late Cretaceous.
Alanqa is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Kem Kem Beds of southeastern Morocco. The name Alanqa comes from the Arabic word العنقاءal-‘Anqā’, for a mythical bird of Arabian culture.
Microtuban is an extinct genus of azhdarchoid pterosaur known from the Late Cretaceous of northern Lebanon.
Xericeps is a genus of pterosaur from the Cenomanian stage of the Late Cretaceous. It was discovered from the Kem Kem Beds of southeastern Morocco.
Afrotapejara is an extinct genus of tapejarid pterosaur discovered in Morocco. The type species, Afrotapejara zouhri, was named and described in 2020. It was the first tapejarid discovered in Africa and the fourth pterosaur discovered in the Kem Kem Beds.
Apatorhamphus is an extinct genus of azhdarchoid pterosaur from the Kem Kem Group of Morocco. It might have been part of the Chaoyangopteridae. It is only known from a few snout fragments and it likely had a wingspan of between 3–7 metres (9.8–23.0 ft)
Nicorhynchus is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, N. capito, from the Cambridge Greensand of England, and N. fluviferox from the Kem Kem Group of Morocco. These species were previously assigned to Coloborhynchus.
Wellnhopterus is an azhdarchid pterosaur recovered from the Late Cretaceous Javelina Formation in Texas that was previously identified as a thalassodromine. It consists of a set of upper and lower jaws, as well as some cervical vertebrae and a fragmentary long bone. In July 2021, the jaws were given the genus name "Javelinadactylus", with the type and only species as "J. sagebieli"; however, this article has now been retracted. In a paper published in December 2021, the complete holotype was independently named Wellnhopterus, with the only species being W. brevirostris. As of 2022, this is the formal name of this pterosaur.
Akharhynchus is an extinct genus of tropeognathine pteranodontoid pterosaurs possibly from the Cretaceous Kem Kem Group of Morocco. The genus contains a single species, A. martilli, known from a small fragment of the premaxillae.