Simurghia

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Simurghia
Temporal range: Late Cretaceous, 66  Ma
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Holotype humerus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Clade: Pteranodontia
Clade: Aponyctosauria
Genus: Simurghia
Longrich, Martill, & Andres, 2018
Type species
Simurghia robusta
Longrich, Martill, & Andres, 2018

Simurghia is a genus of pterosaur from the Ouled Abdoun Basin of Morocco, a basin that dates to the Maastrichtian stage of the Late Cretaceous period, about 66 million years ago. It was published in 2018 by paleontologists Nicholas R. Longrich, David M. Martill, and Brian Andres, along with two other pterosaurs from the same basin: Alcione and Barbaridactylus . [1] The type and only species is S. robusta.

Contents

Discovery and naming

All known specimens of Simurghia were uncovered in a 3-year dig that unearthed about 200 pterosaur specimens. The type specimen, FSAC-OB 7, consists of a nearly complete humerus, lacking only the humeral head and the ulnar crest. [1]

Simurghia is named after a flying beast from Persian mythology known as the Simurgh. The species name, S. robusta, is a Latin word that means "robust." [1]

Classification

Size of Simurghia (top middle) compared to other pterosaurs, birds, and a human Size disparity of late Maastrichtian pterosaurs and birds.svg
Size of Simurghia (top middle) compared to other pterosaurs, birds, and a human

Below is a cladogram showing the results of a phylogenetic analysis first presented by Andres and colleagues in 2014, and updated with additional data by Longrich and colleagues in 2018. In this analysis, they found Simurghia to be the sister taxon of the species Alcione elainus , both placed within the family Nyctosauridae. [2] [1]

  Pteranodontia  
Pteranodontidae

Pteranodon sternbergi

Pteranodon longiceps

Tethydraco regalis

Nyctosauridae

Alamodactylus byrdi

Volgadraco bogolubovi

Cretornis hlavaci

Alcione elainus

Simurghia robusta

Muzquizopteryx coahuilensis

Barbaridactylus grandis

Nyctosaurus lamegoi

Nyctosaurus nanus

Nyctosaurus gracilis

In their phylogenetic analyses, Fernandes et al. (2022) recovered Simurghia as the sister taxon to Epapatelo and Alcione . These taxa, along with the Nyctosauridae, comprise the new clade, Aponyctosauria. [3]

Pteranodontia

Tethydraco

Pteranodon longiceps

Pteranodon sternbergi( Geosternbergia )

Alamodactylus

Volgadraco

Cretornis

Aponyctosauria

Epapatelo

Simurghia

Alcione

Nyctosauridae

Muzquizopteryx

Nyctosaurus lamegoi

Nyctosaurus grandis( Barbaridactylus )

Nyctosaurus nanus

Nyctosaurus gracilis

Paleobiology

Simurghia is similar in form to Alcione , but is unlikely to represent an adult of this species. All specimens of Alcione are either subadults or adults, based on their bone's dense avascular surface texture, well ossified condyles, and fused synsacrum and scapulocoracoid. Additionally, lack of intermediately sized humeri would suggest that these pterosaurs are distinct from each other. Using isometric scaling, Simurghia would weigh 560% more than Alcione, making sexual dimorphism or intraspecific variation unlikely. It is estimated to have a wingspan of 5 m (16 ft). [1]

Paleoecology

Locality and stratigraphy of the discovery site of Simurghia. Phosphate mines in Morocco.PNG
Locality and stratigraphy of the discovery site of Simurghia.

Simurghia was discovered in upper Maastrichtian phosphates located in the Ouled Abdoun Basin of Northern Morocco. This geological formation does not belong to a formation. It is divided into couches, with Simurghia being discovered in Couche III. This layer dates back to the latest Maastrichtian, about 1 million years before the extinction event at the end of the Cretaceous. Couche III represents the most diverse marine ecosystem known from the time. [1]

A large variety of marine life is known from the region, including sharks and bony fish, in addition to turtles, plesiosaurs and mosasaurs. A few rare, fragmentary dinosaur fossils are known, including an abelisaur and a sauropod. Other pterosaurs in the formation were originally only represented by the azhdarchid Phosphatodraco mauritanicus . However, in addition to Simurghia, more recent studies have revealed at least five more pterosaur species from this location, including Barbaridactylus , Alcione and Tethydraco . [1]

Related Research Articles

<span class="mw-page-title-main">Pteranodontidae</span> Family of pteranodontian pterosaurs

The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head. The spectacularly-crested Nyctosaurus is sometimes included in this family, though usually placed in its own family, the Nyctosauridae.

<i>Montanazhdarcho</i> Genus of azhdarchoid pterosaur from the Late Cretaceous

Montanazhdarcho is a genus of azhdarchoid pterosaur from the Late Cretaceous period of what is now the state of Montana, United States. Montanazhdarcho is known from only one species, M. minor.

<i>Phosphatodraco</i> Late Cretaceous genus of pterosaur

Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.

<span class="mw-page-title-main">Lonchodectidae</span> Family of pteranodontoid pterosaurs

Lonchodectidae or Lonchodraconidae is a group of pterosaurs within the clade Pterodactyloidea. It has variously been considered to be within Ctenochasmatoidea, Azhdarchoidea and Pteranodontia. They are notable for their high, conical tooth sockets and raised alveolar margins.

Elanodactylus is a genus of ctenochasmatid pterodactyloid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of Liaoning, China.

<i>Volgadraco</i> Genus of ornithocheiroid pterosaur from the Late Cretaceous

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<span class="mw-page-title-main">Nyctosauridae</span> Family of pteranodontian pterosaurs

Nyctosauridae is a family of specialized soaring pterosaurs of the late Cretaceous Period of North America, Africa, and possibly Europe. It was named in 1889 by Henry Alleyne Nicholson and Richard Lydekker.

<i>Alanqa</i> Genus of azhdarchid pterosaur from the Late Cretaceous

Alanqa is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Kem Kem Beds of southeastern Morocco. The name Alanqa comes from the Arabic word العنقاءal-‘Anqā’, for a mythical bird of Arabian culture.

<span class="mw-page-title-main">Pteranodontia</span> Clade of pteranodontoid pterosaurs

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<span class="mw-page-title-main">Eupterodactyloidea</span> Infraorder of pterodactyloid pterosaurs

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<i>Aerotitan</i> Genus of azhdarchid pterosaur from the Late Cretaceous

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<i>Eurazhdarcho</i> Genus of azhdarchid pterosaur from the Late Cretaceous

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Alamodactylus is an extinct genus of nyctosaurid pteranodontoid pterosaur known from the Late Cretaceous of Texas, southern United States. It contains a single species, Alamodactylus byrdi.

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<span class="mw-page-title-main">Ouled Abdoun Basin</span> Phosphate basin in Morocco

The Oulad Abdoun Basin is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and at least 26.8 billion tons of phosphate. It is also known as an important site for vertebrate fossils, with deposits ranging from the Late Cretaceous (Cenomanian-Turonian) to the Eocene epoch (Ypresian), a period of about 25 million years.

<i>Tethydraco</i> Genus of ornithocheiroid pterosaur from the Late Cretaceous

Tethydraco is a genus of pterodactyloid pterosaur that lived during the Late Cretaceous period of what is now the area of present Morocco, about 66 million years ago. Tethydraco was originally assigned to the family Pteranodontidae. Some researchers argued that subsequently described material suggests that it may have been an azhdarchid, and possibly synonymous with Phosphatodraco, though this has been disputed. The type and only species is T. regalis.

<i>Alcione elainus</i> Genus of pterosaur from the Late Cretaceous

Alcione is a genus of pterosaur from the Ouled Abdoun Basin of Morocco, dating back to the Maastrichtian stage of the Late Cretaceous period. Only one species, A. elainus, is known. This pterosaur lived in a marine environment alongside several other pterosaurs, including Simurghia and Barbaridactylus.

<i>Barbaridactylus</i> Genus of nyctosaurid pterosaur from the Late Cretaceous

Barbaridactylus is a nyctosaurid pterosaur from the Ouled Abdoun Basin of Morocco, a basin that dates back to the Maastrichtian stage of the Late Cretaceous. It was published in 2018 by paleontologists Nicholas R. Longrich, David M. Martill, and Brian Andres. In the same publication, two other pterosaurs from the same basin were described: Alcione and Simurghia. The type and only species is B. grandis.

<i>Thalassotitan</i> Large bodied African mosasaur

Thalassotitan is an extinct genus of large mosasaurs that lived during the late Maastrichtian of the Cretaceous period in what is now Morocco, around 66 million years ago. The only known species is T. atrox, described in 2022 from fossils discovered in the Ouled Abdoun Basin, where many other mosasaurs have been found. It was assigned to the tribe Prognathodontini alongside other mosasaurs like Prognathodon and Gnathomortis. The prognathodontines are separated from other mosasaurs based on their massive jaws and robust teeth.

<i>Epapatelo</i> Genus of pteranodontian pterosaur

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References

  1. 1 2 3 4 5 6 7 Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018). "Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology. 16 (3): e2001663. doi: 10.1371/journal.pbio.2001663 . PMC   5849296 . PMID   29534059.
  2. Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24 (9): 1011–6. doi: 10.1016/j.cub.2014.03.030 . PMID   24768054.
  3. Fernandes, Alexandra E.; Mateus, Octávio; Andres, Brian; Polcyn, Michael J.; Schulp, Anne S.; Gonçalves, António Olímpio; Jacobs, Louis L. (2022). "Pterosaurs from the Late Cretaceous of Angola". Diversity. 14 (9). 741. doi: 10.3390/d14090741 . hdl: 10362/145845 .