Simurghia

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Simurghia
Temporal range: Late Cretaceous, Maastrichtian
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Holotype humerus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Clade: Pteranodontia
Clade: Aponyctosauria
Genus: Simurghia
Longrich, Martill, & Andres, 2018
Type species
Simurghia robusta
Longrich, Martill, & Andres, 2018
Species
  • S. robusta
    Longrich, Martill, & Andres, 2018
  • S. lamegoi? [1]
    (Price, 1953)
Synonyms

Simurghia is a genus of pterosaur from the Ouled Abdoun Basin of Morocco, a basin that dates to the Maastrichtian stage of the Late Cretaceous period, about 66 million years ago. It was published in 2018 by paleontologists Nicholas R. Longrich, David M. Martill, and Brian Andres, along with two other pterosaurs from the same basin: Alcione and Barbaridactylus . [2] The type species is S. robusta. A possible second species, S. ("Nyctosaurus") lamegoi, is known from the Campanian-Maastrichtian age strata of Brazil. [2] [1]

Contents

Discovery and naming

All known specimens of Simurghia were uncovered in a 3-year dig that unearthed about 200 pterosaur specimens. The type specimen, FSAC-OB 7, consists of a nearly complete humerus, lacking only the humeral head and the ulnar crest. [2]

Simurghia is named after a flying beast from Persian mythology known as the Simurgh. The species name, S. robusta, is a Latin word that means "robust." [2]

In 2024, " Nyctosaurus " lamegoi was assigned as a species of Simurghia. [1]

Classification

Size of Simurghia (top middle) compared to other pterosaurs, birds, and a human Size disparity of late Maastrichtian pterosaurs and birds.svg
Size of Simurghia (top middle) compared to other pterosaurs, birds, and a human

Below is a cladogram showing the results of a phylogenetic analysis first presented by Andres and colleagues in 2014, and updated with additional data by Longrich and colleagues in 2018. In this analysis, they found Simurghia to be the sister taxon of the species Alcione elainus , both placed within the family Nyctosauridae. [3] [2]

  Pteranodontia  

In their phylogenetic analyses, Fernandes et al. (2022) recovered Simurghia as the sister taxon to Epapatelo and Alcione . These taxa, along with the Nyctosauridae, comprise the clade Aponyctosauria. [4]

Pteranodontia

Paleobiology

Simurghia is similar in form to Alcione , but is unlikely to represent an adult of this species. All specimens of Alcione are either subadults or adults, based on their bone's dense avascular surface texture, well ossified condyles, and fused synsacrum and scapulocoracoid. Additionally, lack of intermediately sized humeri would suggest that these pterosaurs are distinct from each other. Using isometric scaling, Simurghia would weigh 560% more than Alcione, making sexual dimorphism or intraspecific variation unlikely. It is estimated to have a wingspan of 5 m (16 ft). [2]

Paleoecology

Locality and stratigraphy of the discovery site of Simurghia. Phosphate mines in Morocco.PNG
Locality and stratigraphy of the discovery site of Simurghia.

Simurghia was discovered in upper Maastrichtian phosphates located in the Ouled Abdoun Basin of Northern Morocco. This geological formation does not belong to a formation. It is divided into couches, with Simurghia being discovered in Couche III. This layer dates back to the latest Maastrichtian, about 1 million years before the extinction event at the end of the Cretaceous. Couche III represents the most diverse marine ecosystem known from the time. [2]

A large variety of marine life is known from the region, including sharks and bony fish, in addition to turtles, plesiosaurs and mosasaurs. A few rare, fragmentary dinosaur fossils are known, including an abelisaur and a sauropod. Other pterosaurs in the formation were originally only represented by the azhdarchid Phosphatodraco mauritanicus . However, in addition to Simurghia, more recent studies have revealed at least five more pterosaur species from this location, including Barbaridactylus , Alcione and Tethydraco . [2]

Related Research Articles

<i>Nyctosaurus</i> Genus of nyctosaurid pterosaur from the Late Cretaceous

Nyctosaurus is a genus of nyctosaurid pterosaur from the Late Cretaceous period of what is now the Niobrara Formation of the mid-western United States, which, during the time Nyctosaurus was alive, was covered in an extensive shallow sea. Some remains belonging to a possible Nyctosaurus species called "N." lamegoi have been found in Brazil, though it likely belongs to a different genus Simurghia. The genus Nyctosaurus has had numerous species referred to it, though how many of these may actually be valid requires further study. At least one species possessed an extraordinarily large antler-like cranial crest.

<span class="mw-page-title-main">Pteranodontidae</span> Family of pteranodontian pterosaurs

The Pteranodontidae are a family of large pterosaurs from the Late Cretaceous of North America and possibly other continents including Europe and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head. The spectacularly-crested Nyctosaurus is sometimes included in this family, though usually placed in its own family, the Nyctosauridae.

<i>Phosphatodraco</i> Late Cretaceous genus of pterosaur

Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.

<i>Muzquizopteryx</i> Genus of nyctosaurid pterosaur from the Late Cretaceous

Muzquizopteryx is a genus of nyctosaurid pterodactyloid pterosaur from the Late Cretaceous period of what is now Coahuila, Mexico.

<i>Cretornis</i> Genus of ornithocheiroid pterosaur from the Late Cretaceous

Cretornis is a pterosaur genus from the late Cretaceous period of what is now the Jizera Formation in the Czech Republic, dating to about 92 million years ago. It only contains a single species, Cretornis hlavaci.

Elanodactylus is a genus of ctenochasmatid pterodactyloid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of Liaoning, China.

<i>Volgadraco</i> Genus of ornithocheiroid pterosaur from the Late Cretaceous

Volgadraco is a genus of pteranodontian pterosaur from the Upper Cretaceous of European Russia.

<span class="mw-page-title-main">Nyctosauridae</span> Family of pteranodontian pterosaurs

Nyctosauridae is a family of specialized soaring pterosaurs of the late Cretaceous Period of North America, Africa, and possibly other continents including South America. It was named in 1889 by Henry Alleyne Nicholson and Richard Lydekker.

<i>Alanqa</i> Genus of azhdarchid pterosaur from the Late Cretaceous

Alanqa is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Kem Kem Beds of southeastern Morocco. The name Alanqa comes from the Arabic word العنقاءal-‘Anqā’, for a mythical bird of Arabian culture.

<span class="mw-page-title-main">Pteranodontia</span> Clade of pteranodontoid pterosaurs

Pteranodontia is an extinct group of ornithocheiroid pterodactyloid pterosaurs. It lived during the Late Cretaceous of North America, South America, Europe and Africa. They were some of the most advanced pterosaurs, and possessed highly specialized cranial crests that may have served as sexual attraction, with males having a much larger crest.

<span class="mw-page-title-main">Eupterodactyloidea</span> Infraorder of pterodactyloid pterosaurs

Eupterodactyloidea is an extinct group of pterodactyloid pterosaurs that existed from the latest Late Jurassic to the latest Late Cretaceous periods. Eupterodactyloids have been found on all continents except Antarctica.

<i>Aerotitan</i> Genus of azhdarchid pterosaur from the Late Cretaceous

Aerotitan is a genus of large azhdarchid pterosaur known from the Late Cretaceous period of what is now the Allen Formation of the Neuquén Basin in northern Patagonia, Argentina.

Alamodactylus is an extinct genus of pteranodontian pterosaur known from the Late Cretaceous of Texas, southern United States. It contains a single species, Alamodactylus byrdi.

<span class="mw-page-title-main">Ouled Abdoun Basin</span> Phosphate basin in Morocco

The Oulad Abdoun Basin is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and at least 26.8 billion tons of phosphate. It is also known as an important site for vertebrate fossils, with deposits ranging from the Late Cretaceous (Cenomanian-Turonian) to the Eocene epoch (Ypresian), a period of about 25 million years.

<i>Tethydraco</i> Genus of ornithocheiroid pterosaur from the Late Cretaceous

Tethydraco is a genus of pterodactyloid pterosaur that lived during the Late Cretaceous period of what is now the area of present Morocco, about 66 million years ago. Tethydraco was originally assigned to the family Pteranodontidae. Some researchers argued that subsequently described material suggests that it may have been an azhdarchid, and possibly synonymous with Phosphatodraco, though this has been disputed. The type and only species is T. regalis.

<i>Alcione elainus</i> Genus of pterosaur from the Late Cretaceous

Alcione is a genus of pterosaur from the Ouled Abdoun Basin of Morocco, dating back to the Maastrichtian stage of the Late Cretaceous period. Only one species, A. elainus, is known. This pterosaur lived in a marine environment alongside several other pterosaurs, including Simurghia and Barbaridactylus.

<i>Barbaridactylus</i> Genus of nyctosaurid pterosaur from the Late Cretaceous

Barbaridactylus is a nyctosaurid pterosaur from the Ouled Abdoun Basin of Morocco, a basin that dates back to the Maastrichtian stage of the Late Cretaceous. It was published in 2018 by paleontologists Nicholas R. Longrich, David M. Martill, and Brian Andres. In the same publication, two other pterosaurs from the same basin were described: Alcione and Simurghia. The type and only species is B. grandis.

<i>Gavialimimus</i> Extinct genus of lizards

Gavialimimus is an extinct genus of plioplatecarpine mosasaur from the Maastrichtian of Morocco. The holotype MHNM.KHG.1231, an articulated skull and associated fragmentary postcrania, was found in the Ouled Abdoun Basin.

<i>Thalassotitan</i> Large bodied African mosasaur

Thalassotitan is an extinct genus of large mosasaurs that lived during the late Maastrichtian of the Cretaceous period in what is now Morocco, around 67 to 66 million years ago. The only known species is T. atrox, described in 2022 from fossils discovered in the Ouled Abdoun Basin, initially identified as coming from other genera such as Mosasaurus or Prognathodon. Hypothetical Thalassotitan specimens may have been found in other corners of the world, although researchers also note the possibility that they come from distinct, related taxa. It is considered to be close to the genera Prognathodon and Gnathomortis, together forming the tribe Prognathodontini. The prognathodontines are separated from other mosasaurs based on their massive jaws and robust teeth.

<i>Epapatelo</i> Genus of pteranodontian pterosaur

Epapatelo is a genus of pteranodontian pterosaur from the late Cretaceous Mocuio Formation of Namibe Province, Angola. The genus contains a single species, E. otyikokolo, known from partial left limb bones.

References

  1. 1 2 3 Pêgas, Rodrigo V. (2024-06-10). "A taxonomic note on the tapejarid pterosaurs from the Pterosaur Graveyard site (Caiuá Group, ?Early Cretaceous of Southern Brazil): evidence for the presence of two species". Historical Biology : 1–22. doi:10.1080/08912963.2024.2355664. ISSN   0891-2963.
  2. 1 2 3 4 5 6 7 8 Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018). "Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology. 16 (3): e2001663. doi: 10.1371/journal.pbio.2001663 . PMC   5849296 . PMID   29534059.
  3. Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24 (9): 1011–6. Bibcode:2014CBio...24.1011A. doi: 10.1016/j.cub.2014.03.030 . PMID   24768054.
  4. Fernandes, Alexandra E.; Mateus, Octávio; Andres, Brian; Polcyn, Michael J.; Schulp, Anne S.; Gonçalves, António Olímpio; Jacobs, Louis L. (2022). "Pterosaurs from the Late Cretaceous of Angola". Diversity. 14 (9). 741. doi: 10.3390/d14090741 . hdl: 10362/145845 .