Rhamphorhynchids Temporal range: Early-Late Jurassic, | |
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Fossil specimen of Rhamphorhynchus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Clade: | † Novialoidea |
Clade: | † Breviquartossa Unwin, 2003 |
Family: | † Rhamphorhynchidae Seeley, 1870 |
Type species | |
† Pterodactylus longicaudus Münster, 1839 | |
Subgroups [1] | |
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Synonyms | |
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Rhamphorhynchidae is a group of early pterosaurs named after Rhamphorhynchus , that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley. [10] Members of the group possess no more than 11 pairs of teeth in the rostrum, a deltopectoral crest that is constricted at the base but expanded at the distal end, and a bent phalange on the fifth toe. [11] [12]
Rhamphorhynchidae traditionally contains two subfamilies: the Rhamphorhynchinae and the Scaphognathinae. While not recovered as distinct clades by all analyses, there do appear to be traits uniting members of each group. [1] [12] Rhamphorhynchines are more common, were lightly built, and had jaws ending in pointed tips that contained more teeth, which are often procumbent (pointed forward). Scaphognathines are comparatively quite rare, were more robust skeletally, and had shorter wing proportions. The broad-tipped skulls of scaphognathines held fewer teeth which were widely spaced and vertically oriented in the jaw. [12]
The cladogram (family tree) of rhamphorhynchids below is the result of a large phylogenetic analysis published by Andres & Myers in 2013. [1] Note that this study does not recover Scaphognathinae.
Sordes was a small pterosaur from the late Jurassic (Oxfordian/Kimmeridgian) Karabastau Svita of Kazakhstan.
Anurognathidae is a family of small, short-tailed pterosaurs that lived in Europe, Asia, and possibly North America during the Jurassic and Cretaceous periods. Five genera are known: Anurognathus, from the Late Jurassic of Germany; Jeholopterus, from the Middle to Late Jurassic of China; Dendrorhynchoides, from the Middle Jurassic of China; Batrachognathus, from the Late Jurassic of Kazakhstan; and Vesperopterylus, from the Early Cretaceous of China. Bennett (2007) suggested that the holotype of Mesadactylus, BYU 2024, a synsacrum, belonged to an anurognathid, though this affinity has been questioned by other authors. Mesadactylus is from the Late Jurassic Morrison Formation of the United States. Indeterminate anurognathid remains have also been reported from the Middle Jurassic Bakhar Svita of Mongolia and the Early Cretaceous of North Korea.
The Rhamphorhynchoidea forms one of the two suborders of pterosaurs and represents an evolutionary grade of primitive members of flying reptiles. This suborder is paraphyletic unlike the Pterodactyloidea, which arose from within the Rhamphorhynchoidea as opposed to a more distant common ancestor. Because it is not a completely natural grouping, Rhamphorhynchoidea is not used as a formal group in most scientific literature, though some pterosaur scientists continue to use it as an informal grouping in popular works, such as The Pterosaurs: From Deep Time by David Unwin, and in some formal studies. Rhamphorhynchoids were the first pterosaurs to have appeared, in the late Triassic Period. Unlike their descendants, the pterodactyloids, most rhamphorhynchoids had teeth and long tails, and most species lacked a bony crest, though several are known to have crests formed from soft tissue like keratin. They were generally small, with wingspans rarely exceeding 2.5 meters, though one specimen alluded to by Alexander Stoyanow would be among the largest pterosaurs of all time with a wingspan of 10 meters, comparable to the largest azhdarchids. However, this alleged giant Jurassic pterosaur specimen is not recorded anywhere outside the original Time article. Nearly all rhamphorhynchoids had become extinct by the end of the Jurassic Period, though some anurognathids persisted to the early Cretaceous. The family Wukongopteridae, which shows a mix of rhamphorhynchoid and pterodactyloid features, is known from the Daohugou Beds which are most commonly dated to the Jurassic, but a few studies give a Cretaceous date. Furthermore, remains of a non-pterodactyloid from the Candeleros Formation extend the presence of basal pterosaurs into at least the early Late Cretaceous.
Boreopterus is a genus of boreopterid pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Dalian, Liaoning, China.
Dendrorhynchoides was a genus of anurognathid pterosaur containing only the holotype species D. curvidentatus that is known from the Middle Jurassic Tiaojishan Formation of Qinglong, northern Hebei Province, China.
Eoazhdarcho is a genus of azhdarchoid pterodactyloid pterosaur named in 2005 by Chinese paleontologists Lü Junchang and Ji Qiang. The type and only known species is Eoazhdarcho liaoxiensis. The fossil was found in the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China.
Parapsicephalus is a genus of long-tailed rhamphorhynchid pterosaurs from the Lower Jurassic Whitby, Yorkshire, England. It contains a single species, P. purdoni, named initially as a species of the related rhamphorhynchid Scaphognathus in 1888 but moved to its own genus in 1919 on account of a unique combination of characteristics. In particular, the top surface of the skull of Parapsicephalus is convex, which is otherwise only seen in dimorphodontians. This has been the basis of its referral to the Dimorphodontia by some researchers, but it is generally agreed upon that Parapsicephalus probably represents a rhamphorhynchid. Within the Rhamphorhynchidae, Parapsicephalus has been synonymized with the roughly contemporary Dorygnathus; this, however, is not likely given the many differences between the two taxa, including the aforementioned convex top surface of the skull. Parapsicephalus has been tentatively referred to the Rhamphorhynchinae subgrouping of rhamphorhynchids, but it may represent a basal member of the group instead.
Liaoxipterus is a genus of pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. The type species is Liaoxipterus brachyognathus. The genus name is derived from the discovery locality Liaoxi and a Latinised Greek pteron, "wing". The specific name is derived from Greek brachys, "short" and gnathos, "jaw".
The Tuchengzi Formation is a geological formation in China whose strata span the Tithonian to Berriasian ages. Dinosaur fossils, particularly footprints, have been found from the formation.
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period. It is known for its exceptionally preserved fossils, including those of plants, insects and vertebrates. It is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks. Previously, the Tiaojishan Formation was grouped together with the underlying Haifanggou Formation as a single "Lanqi Formation." The Tiaojishan Formation forms a key part of the Yanliao Biota assemblage, alongside the Haifanggou Formation.
Zhenyuanopterus is a genus of boreopterid pterosaur which is known from Lower Cretaceous Yixian Formation of Liaoning, China. It contains one species, Zhenyuanopterus longirostris, which was first described and named by Lü Junchang.
Fenghuangopterus is a genus of basal pterosaur that lived in northeastern China during the Middle Jurassic.
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of Pterodactylus and Ctenochasma, and all their descendants.
Boreopteridae is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China.
Jianchangopterus is a genus of scaphognathine rhamphorhynchid pterosaur from the Middle Jurassic of western Liaoning, China. Jianchangopterus is known from a nearly complete skeleton with skull preserved. It was collected from the Tiaojishan Formation. It was first named by Lü Junchang and Bo Xue in 2011 and the type species is Jianchangopterus zhaoianus.
Qinglongopterus is a genus of rhamphorhynchid pterosaur from the Middle/Upper Jurassic of Mutoudeng, Qinglong County, Hebei Province, China. Qinglongopterus is known from only one specimen; D3080/3081, a nearly complete skeleton collected from the Tiaojishan Formation. It was described by Lü Junchang et al. in 2012. The type species is Qinglongopterus guoi.
This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs went extinct millions of years before humans evolved, humans have coexisted with pterosaur fossils for millennia. Before the development of paleontology as a formal science, these remains would have been interpreted through a mythological lens. Myths about thunderbirds told by the Native Americans of the modern Western United States may have been influenced by observations of Pteranodon fossils. These thunderbirds were said to have warred with water monsters, which agrees well with the co-occurrence of Pteranodon and the ancient marine reptiles of the seaway over which it flew.
Orientognathus is a genus of rhamphorhynchid pterosaur from the Middle Jurassic of China. It is known from a single specimen which includes most of the skeleton and skull, and was first named and described in 2015 by Lü Junchang et al.. The taxon was found in the Tuchengzi Formation of China, which is slightly younger than the Tiaojishan Formation that most other Middle Jurassic pterosaurs from the region have been found in. The description study produced a phylogenetic analysis, which determined that Orientognathus was a basal member of Rhamphorhynchidae, possibly within Rhamphorhynchinae.