Lonchodectes Temporal range: Late Cretaceous, | |
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Lectotype jaw fragment (A–D) and assigned rostrum fragment (E–H) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Ornithocheiromorpha |
Family: | † Lonchodectidae Hooley, 1914 |
Genus: | † Lonchodectes Hooley, 1914 |
Type species | |
†Pterodactylus compressirostris Owen, 1851 | |
Species | |
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Synonyms | |
Genus synonymy
Species synonymy
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Lonchodectes (meaning "lance biter") was a genus of lonchodectid pterosaur from several formations dating to the Turonian (Late Cretaceous) of England, mostly in the area around Kent. The species belonging to it had been assigned to Ornithocheirus until David Unwin's work of the 1990s and 2000s. [1] Several potential species are known; most are based on scrappy remains, and have gone through several other generic assignments. The genus is part of the complex taxonomy issues surrounding Early Cretaceous pterosaurs from Brazil and England, such as Amblydectes , Anhanguera , Coloborhynchus , and Ornithocheirus. [2]
Numerous species have been referred to this genus over time, and only those more widely connected with the genus are included here.
The type species, L. compressirostris, is based on NHMUK 39410, a partial upper jaw from the Turonian-age Upper Cretaceous Upper Chalk near Kent. Richard Owen named in 1851 as a species of Pterodactylus ; [3] it was transferred to Ornithocheirus in 1870 by Harry Govier Seeley, [4] before becoming the type species of Lonchodectes in Reginald Walter Hooley's 1914 review of Ornithocheirus. [5] Confusingly, this species was also long regarded, incorrectly, as the type species of Ornithocheirus. [6]
A variety of postcranial remains resembling those of Azhdarchoids from the Cambridge Greensand have been referred to Lonchodectes; [6] [7] [8] however, much of this material has since been referred to Ornithostoma . [9]
In 2019, the Brazilian palaeontologist Rodrigo V. Pêgas and colleagues suggested that the type specimen of Lonchodectes compressirostris could represent the same species as Cimoliopterus cuvieri . They cautioned this is impossible to confirm until associated skull and mandible material is found. [10]
Hooley added two other species at this time, both of which had also been originally referred to Pterodactylus, then to Ornithocheirus: L. giganteus, a Cenomanian-age jaw fragment from the Chalk of Kent; [11] and L. daviesii, another jaw fragment, from the Albian-age Gault Clay. [12]
"Pterodactylus" sagittirostris, based on NHMUK R.1823, a lower jaw fragment from the ?Valanginian-Hauterivian-age Lower Cretaceous Hastings Beds of East Sussex, [12] "Ornithocheirus" platystomus, [4] "Ornithocheirus" machaerorhynchus, and "O." microdon were assigned to Lonchodectes in a 2001 review by David Unwin of Cambridge Greensand pterosaurs. [6] joining L. compressirostris, L. giganteus, L. platystomus, and L. sagittirostris in his listing of valid species. [13] However, L. giganteus, L. machaerorhynchus, and L. microdon have since been assigned to a new genus, Lonchodraco , while L. sagittirostris has been renamed Serradraco . [2] [14] L. platystomus may be a species of Amblydectes . [2] In 2020, a review of Lonchodectidae was conducted by paleontologist Alexander Averianov, where he reassigned the species L. machaerorhynchus to the genus Ikrandraco due to similarities in rostral morphology, as I. machaerorhynchus, and he also considered L. microdon a junior synonym of machaerorhynchus. Therefore, Lonchodectes is limited to its type species, L. compressirostris. [15]
In Peter Wellnhofer's 1991 The Illustrated Encyclopedia of Pterosaurs, written before Unwin's work, the species were included within Ornithocheirus (because of L. compressirostris being thought to be the type species), and are in fact the main fossils illustrated to represent the genus. [16] In 2003, Unwin placed them in their own family, Lonchodectidae, which he grouped within the group Ctenochasmatoidea, [17] while in 2006, he placed the family Lonchodectidae within the Azhdarchoidea, the group that includes the tapejarids and azhdarchids. [13]
The cladogram below is a topology recovered by Longrich and colleagues in 2018. In their analysis, they placed Lonchodectes within the family Lonchodectidae as the sister taxon of Lonchodraco. Contrary to previous analyses, Longrich and colleagues placed Lonchodectidae (including Lonchodectes) within the more inclusive group Ornithocheiromorpha. [18]
Lonchodectes had long jaws with many short teeth, and the jaws were compressed vertically, like "a pair of sugar tongs with teeth". [19] Related species (including several taxa formerly included within the genus) had crests on their lower jaws, so the same probably also applied to L. compressirostris. [20]
Pterodactylus is a genus of extinct pterosaurs. It is thought to contain only a single species, Pterodactylus antiquus, which was the first pterosaur to be named and identified as a flying reptile and one of the first prehistoric reptiles to ever be discovered.
Ornithocheirus is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the United Kingdom and possibly Morocco.
Tropeognathus is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. Tropeognathus is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is Tropeognathus mesembrinus. Fossil remains of Tropeognathus have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil.
Amblydectes is a genus of pterosaur known from jaw fragments. It apparently had a jaw flattened towards the tip and triangular in cross-section. It has at times been synonymized with Coloborhynchus, Criorhynchus, Lonchodectes, or Ornithocheirus. A 2013 study found A. crassidens and A. eurygnathus to be nomina dubia, with A. platystomus possibly belonging to a separate, yet unnamed genus. A 2021 study found A. crassidens to be a valid genus within Anhangueridae, while A. platystomus was placed in the new genus Draigwenia. A. eurygnathus was found to possibly be a junior synonym of A. crassidens.
Ornithostoma is a genus of pterodactyloid pterosaur that lived during the Early Cretaceous period of Europe, around 110 million years ago. Ornithostoma was once thought to have been a senior synonym of the pteranodontid Pteranodon due to its toothless anatomy and prior naming.
Siroccopteryx is an extinct genus of anhanguerid pterodactyloid pterosaur which lived in Morocco during the Cenomanian stage of the Late Cretaceous. Some researchers, such as David M. Unwin, consider the genus a junior synonym of Coloborhynchus.
Coloborhynchus is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. Coloborhynchus is known from the Lower Cretaceous of England, and depending on which species are included, possibly the Albian and Cenomanian ages as well. Coloborhynchus was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related Tropeognathus is now thought to have had a larger wingspan.
Lonchodectidae or Lonchodraconidae is a group of pterosaurs within the clade Pterodactyloidea. It has variously been considered to be within Ctenochasmatoidea, Azhdarchoidea and Pteranodontoidea. They are notable for their high, conical tooth sockets and raised alveolar margins.
"Palaeornis" cliftii is a pterosaur species known from parts of a single humerus found in the early Cretaceous (Valanginian) of the upper Tunbridge Wells Sand Formation, England.
Ornithocheiridae is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cretaceous periods, around 140 to 90 million years ago.
The Cambridge Greensand is a geological unit in England whose strata are earliest Cenomanian in age. It lies above the erosive contact between the Gault Formation and the Chalk Group in the vicinity of Cambridgeshire, and technically forms the lowest member bed of the West Melbury Marly Chalk Formation. It is a remanié deposit, containing reworked fossils of late Albian age, including those of dinosaurs and pterosaurs.
Pteranodontoidea is an extinct clade of ornithocheiroid pterosaurs from the Early to Late Cretaceous of Asia, Africa, Europe, North America and South America. It was named by Alexander Wilhelm Armin Kellner in 1996. In 2003, Kellner defined the clade as a node-based taxon consisting of the last common ancestor of Anhanguera, Pteranodon and all its descendants. The clade Ornithocheiroidea is sometimes considered to be the senior synonym of Pteranodontoidea, however it depends on its definition. Brian Andres in his analyses, converts Ornithocheiroidea using the definition of Kellner (2003) to avoid this synonymy.
Cimoliopterus is a genus of pterosaur that lived during the Late Cretaceous in what is now England and the United States. The first known specimen, consisting of the front part of a snout including part of a crest, was discovered in the Grey Chalk Subgroup of Kent, England, and described as the new species Pterodactylus cuvieri in 1851. The specific name cuvieri honours the palaeontologist George Cuvier, whereas the genus Pterodactylus was then used for many pterosaur species that are not thought to be closely related today. It was among the first pterosaurs to be depicted as sculptures, in Crystal Palace Park in the 1850s. The species was subsequently assigned to various other genera, including Ornithocheirus and Anhanguera. In 2013, the species was moved to a new genus, as Cimoliopterus cuvieri; the generic name Cimoliopterus is derived from the Greek words for "chalk" and "wing". Other specimens and species have also been assigned to or synonymised with the species with various levels of certainty. In 2015, a snout discovered in the Britton Formation of Texas, US, was named as a new species in the genus, C. dunni; the specific name honours its collector, Brent Dunn.
Lonchodraco is a genus of lonchodraconid pterodactyloid pterosaur from the Late Cretaceous of southern England. The genus includes species that were previously assigned to other genera.
Camposipterus is a genus of pterodactyloid pterosaur from the Early Cretaceous of England. Fossil remains of Camposipterus dated back to the Early Cretaceous, about 112 million years ago.
Ikrandraco is a genus of lonchodraconid pterodactyloid pterosaur known from the Lower Cretaceous Jiufotang Formation of northeastern China, and the Cambridge Greensand of England. It is notable for its unusual skull, which features a crest on the lower jaw.
Ornithocheiromorpha is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods, around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by pteranodontians and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing Ornithocheirus, but not Pteranodon.
Serradraco is a genus of Early Cretaceous pterodactyloid pterosaur from the Valanginian aged Tunbridge Wells Sand Formation in England. Named by Rigal et al. in 2018 with the description of a second specimen, it contains a single species, S. sagittirostris, which was formerly considered a species of Lonchodectes, L. sagittirostris. In 2020, Averianov suggested it did not belong in Lonchodectidae.
Nicorhynchus is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, N. capito, from the Cambridge Greensand of England, and N. fluviferox from the Kem Kem Group of Morocco. These species were previously assigned to Coloborhynchus.
Draigwenia is an extinct genus of pterosaur known from a jaw fragment found in the Late Cretaceous Cambridge Greensand in the United Kingdom. The fossil was likely reworked from an Early Cretaceous layer that can be dated to the Albian. It currently contains a single species, Draigwenia platystomus.