Cryodrakon Temporal range: Campanian, | |
---|---|
Life restoration | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Family: | † Azhdarchidae |
Subfamily: | † Quetzalcoatlinae |
Genus: | † Cryodrakon Hone et al., 2019 [1] |
Species: | †C. boreas |
Binomial name | |
†Cryodrakon boreas Hone et al., 2019 [1] | |
Cryodrakon ("cold dragon") is a genus of azhdarchid pterosaur that lived during the Late Cretaceous period in what is now Canada. It contains a single species, Cryodrakon boreas, recovered from the Dinosaur Park Formation. [1] [2] [3]
Since 1972, bones of large azhdarchids had been reported from Alberta, the first pterosaur fossils discovered from Canada. [4] These were sometimes referred to a Quetzalcoatlus sp. [5] Later research by Michael Habib indicated that they represented a taxon new to science. [2]
In 2019, the type species Cryodrakon boreas was named and described by Habib alongside David William Elliott Hone and François Therrien. The generic name is derived from the Greek κρύος (kryos, "icy cold") and δράκων (drakon, "dragon"). The specific name means "belonging to Boreas , the north wind", or simply "boreal" or "northern". [1] Habib had previously considered the name Cryodrakon viserion , as a reference to the ice dragon in Game of Thrones . [2]
The holotype, TMP 1992.83, was found in a layer of the Dinosaur Park Formation dating from the late Campanian (between 76.7 and 74.3 mya), with the excavation site being located near the lower or older boundary of this range. The specimen consists of a partial skeleton, albeit lacking the skull, but contains a fourth cervical vertebra, a rib, a humerus, a pteroid bone, a fourth metacarpal, a tibia and a metatarsal. [2] In 1992, the skeleton was excavated in Quarry Q207; in 1995, it was reported and partially described. [6] The bones were not articulated, but associated, representing an immature animal. It is the only pterosaur skeleton, as opposed to single bones, ever found in Canada. It was described in more detail in 2005. [7]
All the known azhdarchid material from the Dinosaur Park Formation was referred to the species. The specimens consisted of the neck vertebrae TMP 1996.12.369, TMP 1981.16.107, TMP 1980.16.1367, TMP 1989.36.254 and TMP 1993.40.11; the scapulocoracoid TMP 1981.16.182; the ulna TMP 1965.14.398; fourth metacarpals TMP 1979.14.24, TMP 1987.36.16 and TMP 2005.12.156; wing finger phalanges TMP 1972.1.1, TMP 1982.19.295 and TMP 1992.36.936; and the thighbone TMP 1988.36.92. The bones represent individuals of various biological ages, among them juveniles and a large mature exemplar. Most bones are from medium-sized animals. In 2019, only the neck vertebrae were described in detail; bones from other parts of the body had already been treated in 2005. [2]
The material indicates individuals of varying size. Specimen TMP 1996.12.369, a fifth neck vertebra with a length of only 10.6 millimeters (0.42 in), is from a juvenile animal with an estimated wingspan of about 2 meters (6.6 ft). [1] Most bones, among them the holotype, are similar in dimensions to those of Quetzalcoatlus lawsoni, [1] implying a wingspan of about 5 meters (16 ft). [2] Specimen TMP 1980.16.1367, originally mistaken as a leg bone in 1982, is a fifth neck vertebra with an estimated original length of 50 centimeters (1.6 ft), indicating an animal equal in size to the holotype of Quetzalcoatlus northropi, [1] the wingspan of which has been initially estimated at 13 meters (43 ft), [5] but then moderated at 10 meters (33 ft). [1] [2] [8]
Cryodrakon was proportionally similar to Quetzalcoatlus and other long-necked advanced azhdarchids, though its somewhat more robust bones may indicate that it was slightly heavier. [1]
Cryodrakon is distinguished from all other known azhdarchids by two features of its neck vertebrae. The lateral pneumatic fossae or pneumatophores, a pair of small openings leading to air pockets on either side of the neural canal, were positioned near the lower edge of the neural canal, while those of other azhdarchids (with the purported exception of Eurazhdarcho ) were positioned higher up. The second distinguishing feature related to its postexapophyses, large bony knobs adjacent to the protruding rear connection surface of each vertebra, the cotyle. Cryodrakon's postexapophyses were prominent in width but short in length, clearly separated from the cotyle, and their facets were directed downwards. [1]
Cryodrakon was placed in the Azhdarchidae in 2019. No exact cladistic analysis was given clarifying the precise relationships with other azhdarchids. It would have been one of the oldest known azhdarchids of North-America. [1] Then in 2021, Brian Andres performed a phylogenetic analysis on Quetzalcoatlus , including Cryodrakon in the study. It was in a trichotomy with Wellnhopterus brevirostris and Quetzalcoatlus northropi, Q. lawsoni, Hatzegopteryx thambena , and Arambourgiania philadelphiae . Below are the results of the study. [9]
Cryodrakon is thought to have been capable of flight, [2] and if it was, it would have been one of the largest flying animals known to have existed. [3] Azhdarchid pterosaurs similar to Cryodrakon are currently thought to have fed by hunting for smaller animals while on the ground, similar to modern day marabou storks. [10]
The Dinosaur Park Formation, where fossils of Cryodrakon have been excavated, preserves many fossils from alluvial and coastal plain environments. Both of these environments would have experienced high precipitation, warm temperatures, and high humidity, with conditions becoming more swamp-like as time progressed. [11] There was a great diversity of herbivorous dinosaurs including both lambeosaurine and saurolophine hadrosaurs, centrosaurine and chasmosaurine ceratopsians, and nodosaurid and ankylosaurid ankylosaurs, which could have been able to feed on different vegetation levels from one another. [7] [12] The predatory niches were occupied by small maniraptoran theropods, medium-sized juvenile tyrannosaurids, and fully grown tyrannosaurids. [13]
A partial skeleton was found to have been scavenged by a dromaeosaurid, possibly Saurornitholestes langstoni . A broken tooth was found in one of the bones, which according to the authors, meant that the thin-walled bone "must have been very tough." [6]
Albertosaurus is a genus of large tyrannosaurid theropod dinosaur that lived in northwestern North America during the early to middle Maastrichtian age of the Late Cretaceous period, about 71 million years ago. The type species, A. sarcophagus, was apparently restricted in range to the modern-day Canadian province of Alberta, after which the genus is named, although an indeterminate species has been discovered in the Corral de Enmedio and Packard Formations of Mexico. Scientists disagree on the content of the genus and some recognize Gorgosaurus libratus as a second species.
Quetzalcoatlus is a genus of azhdarchid pterosaur known from the Late Cretaceous Maastrichtian age of North America. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is Q. northropi, named by Douglas Lawson in 1975 after the tailless fixed-wing aircraft designer Jack Northrop. The genus also includes the smaller species Q. lawsoni, which was known for many years as an unnamed species, before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Q. northropi has gained fame as a candidate for the largest flying animal ever discovered.
Daspletosaurus is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 78 and 74.4 million years ago, during the Late Cretaceous Period. The genus Daspletosaurus contains three named species. Fossils of the earlier type species, D. torosus, have been found in Alberta, while fossils of a later species, D. horneri, have been found only in Montana. D. wilsoni has been suggested as an intermediate species between D. torosus and D. horneri that evolved through anagenesis, but this theory has been disputed by other researchers.
Gorgosaurus is a genus of tyrannosaurid theropod dinosaur that lived in western North America during the Late Cretaceous Period (Campanian), between about 76.6 and 75.1 million years ago. Fossil remains have been found in the Canadian province of Alberta and the U.S. state of Montana. Paleontologists recognize only the type species, G. libratus, although other species have been erroneously referred to the genus.
Saurornitholestes is a genus of carnivorous dromaeosaurid theropod dinosaur from the late Cretaceous of Canada (Alberta) and the United States.
Philip John Currie is a Canadian palaeontologist and museum curator who helped found the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta and is now a professor at the University of Alberta in Edmonton. In the 1980s, he became the director of the Canada-China Dinosaur Project, the first cooperative palaeontological partnering between China and the West since the Central Asiatic Expeditions in the 1920s, and helped describe some of the first feathered dinosaurs. He is one of the primary editors of the influential Encyclopedia of Dinosaurs, and his areas of expertise include theropods, the origin of birds, and dinosaurian migration patterns and herding behavior. He was one of the models for palaeontologist Alan Grant in the film Jurassic Park.
Azhdarchidae is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well. Azhdarchids are mainly known for including some of the largest flying animals discovered, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and Titanopteryx. They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. Previously it was thought that by the end of the Cretaceous, most pterosaur families except for the Azhdarchidae disappeared from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms. In several analyses, some taxa such as Navajodactylus, Bakonydraco and Montanazhdarcho were moved from Azhdarchidae to other clades.
Azhdarcho is a genus of azhdarchid pterosaur from the late Cretaceous Period of the Bissekty Formation of Uzbekistan, as well as the Zhirkindek Formation of Kazakhstan and possibly also the Ialovachsk Formation of Tajikistan. It is known from fragmentary remains including the distinctive, elongated neck vertebrae that characterizes members of the family Azhdarchidae, a family that includes many giant pterosaurs such as Quetzalcoatlus. The name Azhdarcho comes from the Persian word azhdar (اژدر), a dragon-like creature in Persian mythology. The type species is Azhdarcho lancicollis. The specific epithet lancicollis is derived from the Latin words lancea and collum ("neck").
Arambourgiania is an extinct genus of azhdarchid pterosaur from the Late Cretaceous period of Jordan, and possibly the United States. Arambourgiania was among the largest members of its family, the Azhdarchidae, and it is also one of the largest flying animals ever known. The incomplete left ulna of the "Sidi Chennane azhdarchid" from Morocco may have also belonged to Arambourgiania.
Hatzegopteryx is a genus of azhdarchid pterosaur found in the late Maastrichtian deposits of the Densuş Ciula Formation, an outcropping in Transylvania, Romania. It is known only from the type species, Hatzegopteryx thambema, named by Buffetaut et al. in 2002 based on parts of the skull and humerus. Additional specimens, including a neck vertebra, were later placed in the genus, representing a range of sizes. The largest of these remains indicate it was among the biggest pterosaurs, with an estimated wingspan of 10 to 12 metres.
Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.
Pterosaurs included the largest flying animals ever to have lived. They are a clade of prehistoric archosaurian reptiles closely related to dinosaurs. Species among pterosaurs occupied several types of environments, which ranged from aquatic to forested. Below are the lists that comprise the smallest and the largest pterosaurs known as of 2022.
Alanqa is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Kem Kem Beds of southeastern Morocco. The name Alanqa comes from the Arabic word العنقاءal-‘Anqā’, for a mythical bird of Arabian culture.
Continuoolithus is an oogenus of dinosaur egg found in the late Cretaceous of North America. It is most commonly known from the late Campanian of Alberta and Montana, but specimens have also been found dating to the older Santonian and the younger Maastrichtian. It was laid by an unknown type of theropod. These small eggs are similar to the eggs of oviraptorid dinosaurs, but have a distinctive type of ornamentation.
Navajodactylus is an extinct genus of pterodactyloid pterosaur from Late Cretaceous deposits of the San Juan Basin in New Mexico, United States.
Eurazhdarcho is a genus of azhdarchid pterosaur from the Late Cretaceous period of what is now the Transylvanian Basin of Romania. Its fossil remains dated back 69 million years ago.
Wendy Sloboda is a Canadian fossil hunter from Warner, Alberta. She has made fossil discoveries of dinosaurs and other extinct animals on several continents, with finds in Canada, Argentina, Mongolia, France, and Greenland. She is commemorated in name of the horned dinosaur Wendiceratops, remains of which she discovered in 2010, as well as the fossil footprint Barrosopus slobodai which she discovered in 2003.
Rativates is a genus of ornithomimid theropod dinosaur from the Dinosaur Park Formation of Alberta. The type species is Rativates evadens.
Citipes is an extinct genus of caenagnathid theropod from the Late Cretaceous Dinosaur Park Formation in Alberta, Canada. The genus contains only one species, the type species, C. elegans. The generic name of Citipes is Latin for "fleet-footed", and the specific epithet "elegans" is Latin for "elegant". The type specimen of Citipes has a convoluted taxonomic history, and has been previously assigned to the genera Ornithomimus, Macrophalangia, Elmisaurus, Chirostenotes, and Leptorhynchos before being given its own genus in 2020.
Thanatosdrakon is a genus of quetzalcoatline azhdarchid pterosaur from the Late Cretaceous Plottier Formation of the Neuquén Basin in western Argentina. The genus name is derived from the Greek words thanatos (=death) and drakon (=dragon), while the specific name is a Quechuan word meaning "flying serpent" and refers to the Incan deity Amaru. The type and only species is Thanatosdrakon amaru, known from two specimens consisting of several well-preserved axial and appendicular bones including material previously undescribed in giant azhdarchids. Thanatosdrakon is one of the oldest known members of the Quetzalcoatlinae. T. amaru lived from about 90 to 86 million years ago.