Nipponopterus Temporal range: Late Cretaceous, | |
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Skeletal reconstruction | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Family: | † Azhdarchidae |
Subfamily: | † Quetzalcoatlinae |
Genus: | † Nipponopterus Zhou et al., 2024 |
Species: | †N. mifunensis |
Binomial name | |
†Nipponopterus mifunensis Zhou et al., 2024 | |
Nipponopterus (meaning "Nippon wing") is an extinct genus of azhdarchid pterosaurs from the Late Cretaceous Mifune Group of Japan. The genus contains a single species, N. mifunensis, known from a partial neck vertebra. Nipponopterus represents the first pterosaur to be named from Japan.
The Nipponopterus holotype specimen, MDM 349, was discovered in sediments of the "Upper Formation" of the Mifune Group ('Locality 1018') in rock outcrops near Amagimi Dam in Mifune Town, Kumamoto Prefecture in Kyūshū, Japan. The specimen consists of the posterior end of the sixth cervical vertebra. The specimen was first described in 2000 as belonging to an indeterminate azhdarchid. The limited understanding of the clade at that time precluded more detailed analyses. [1] While early reviews interpreted the cervical vertebra as the fourth [2] or fifth, [3] the most recent analysis suggests it can confidently be regarded as the sixth. [4]
In 2024, Zhou et al. described Nipponopterus mifunensis as a new genus and species of azhdarchid pterosaurs based on these fossil remains. The generic name, Nipponopterus, combines "Nippon"—the Japanese name for the country—with "pterus", derived from the Ancient Greek word ptéron, meaning "wing". The specific name, mifunensis, references the type locality in the Mifune Group in Mifune Town. Nipponopterus is the first pterosaur named from Japan. [4]
A wing phalanx and metacarpal belonging to indeterminate azhdarchid pterosaurs are also known from this formation, but have not been referred to Nipponopterus. [4]
The Nipponopterus holotype is morphologically comparable to a similarly fragmentary unnamed pterosaur specimen from Mongolia, nicknamed the "Burkhant azhdarchid". [5] Measurements of the Nipponopterus material indicates that it is 82% the size of the Burkhant specimen. However, the former likely belongs to a subadult individual, so was not fully grown. As such, Zhou et al. (2024) estimate that both pterosaurs would have had similar wingspans, at around 3–3.5 metres (9.8–11.5 ft). [4]
In their phylogenetic analysis using the dataset of Pêgas (2024), [6] Zhou et al. (2024) recovered Nipponopterus as a member of the azhdarchid clade Quetzalcoatlinae, as the sister taxon to the Burkhant azhdarchid. Their results are displayed in the cladogram below: [4]
Azhdarchomorpha |
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Quetzalcoatlus is a genus of azhdarchid pterosaur known from the Late Cretaceous Maastrichtian age of North America. The first specimen, recovered in 1971 from the Javelina Formation, consists of several wing fragments. It was made the holotype of Quetzalcoatlus northropi in 1975 by Douglas Lawson and was named after the Aztec serpent god, Quetzalcōātl, and Jack Northrop, designer of tailless fixed-wing aircraft. The remains of a second species were found between 1972 and 1974, also by Lawson, around 40 km (25 mi) from the type of Q. northropi. In 2021, these remains were assigned the name Quetzalcoatlus lawsoni by Brian Andres and (posthumously) Wann Langston Jr.
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Aerotitan is a genus of large azhdarchid pterosaur known from the Late Cretaceous period of what is now the Allen Formation of the Neuquén Basin in northern Patagonia, Argentina.
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