Albadraco

Last updated

Albadraco
Temporal range: Maastrichtian, 66  Ma
O
S
D
C
P
T
J
K
Pg
N
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Family: Azhdarchidae
Genus: Albadraco
Solomon, Codrea, Venczel & Grellet-Tinner, 2019
Type species
Albadraco tharmisensis
Solomon, Codrea, Venczel & Grellet-Tinner, 2019

Albadraco is an azhdarchid pterosaur genus that during the Late Cretaceous lived in the area of modern Romania. The type species is Albadraco tharmisensis.

Contents

Discovery and naming

At the site of Oarda de Jos, near Alba Iulia in Alba County, two jaw pieces were found of a large pterosaur. The discovery was reported and illustrated in a dissertation by Cătălin Jipa-Murzea in 2012. [1]

In 2019, the type species Albadraco tharmisensis was named and described by Alexandru Adrian Solomon, Vlad Aurel Codrea, Márton Venczel and Gerald Grellet-Tinner. The generic name combines a reference to Alba with the Latin draco, "dragon". The specific name refers to a provenance near Tharmis, the ancient Dacian name of Alba Iulia. [2]

The holotype, PSMUBB V651a, b, was found in a layer of the Șard Formation dating from the latest Maastrichtian, sixty-six million years old. It consists of two fused premaxillae of the snout (PSMUBB V651a) and a piece of the symphysis of the lower jaws (PSMUBB V651b). Both parts were assumed to have belonged to a single individual, perhaps a subadult animal. It is the first example from the Cretaceous of Europe of a pterosaur exemplar preserving both upper jaw and lower jaw elements. [2]

A second specimen, PSMUBB V652, a fourth neck vertebra from the same site, was referred to the species. It too is from a subadult animal and the describing authors considered it possible that it represented the same individual as the holotype. [2]

Description

The wingspan of the holotype was estimated at five to six metres, by extrapolation from the jaw pieces. The adult size was estimated at a span of six to seven metres. Albadraco shared its habitat with Hatzegopteryx and the possibility was considered that its holotype was only a juvenile exemplar of that giant pterosaur. This was deemed improbable however, because the holotype bone structure resembled that of subadult animals, not of fast growing young individuals. [2]

The describing authors indicated five distinguishing traits. They were autapomorphies, unique derived characters, in which Albadraco differs from all other known Azhdarchidae. The cutting edges and sides of the beak show a high foramina density. The premaxilla has split-like foramina on the lower and side surfaces but also two rows of foramina on the side. The snout has a triangular cross-section but its top edge is more rounded than with other azhdarchids. The symphysis of the lower jaws has a U-shaped cross-section in front but a V-shaped one at the rear. The fourth neck vertebra has an Elongation Ratio, horizontal length divided by transverse width, that more approaches the ER of the third neck vertebra of other azhdarchids in being shorter than usual. [2]

Phylogeny

In 2019, Albadraco was placed in the Azhdarchidae, based on the comparative method. [2]

See also

Related Research Articles

<i>Quetzalcoatlus</i> Genus of azhdarchid pterosaurs from the Late Cretaceous

Quetzalcoatlus is a genus of azhdarchid pterosaur known from the Late Cretaceous Maastrichtian age of North America. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is Q. northropi, named by Douglas Lawson in 1975. The genus also includes the smaller species Q. lawsoni, which was known for many years as an unnamed species, before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Q. northropi has gained fame as a candidate for the largest flying animal ever discovered.

<i>Magyarosaurus</i> Extinct genus of dinosaurs

Magyarosaurus is a genus of dwarf sauropod dinosaur from late Cretaceous Period in Romania. It is one of the smallest-known adult sauropods, measuring only 6 m (20 ft) in length and 750–1,000 kg (1,650–2,200 lb) in body mass. The type and only certain species is Magyarosaurus dacus. It has been found to be a close relative of Rapetosaurus in the family Saltasauridae in the sauropod clade Titanosauria in a 2005 study.

<span class="mw-page-title-main">Azhdarchidae</span> Family of large azhdarchoid pterosaurs

Azhdarchidae is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well. Azhdarchids are mainly known for including some of the largest flying animals discovered, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and Titanopteryx. They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. Previously it was thought that by the end of the Cretaceous, most pterosaur families except for the Azhdarchidae disappeared from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms. In several analyses, some taxa such as Navajodactylus, Bakonydraco and Montanazhdarcho were moved from Azhdarchidae to other clades.

<i>Thalassodromeus</i> Genus of azhdarchoid pterosaur from the Early Cretaceous

Thalassodromeus is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of Thalassodromeus sethi by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner", and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to T. sethi in 2005, became the basis of the new genus Banguela in 2014, and assigned back to Thalassodromeus as the species T. oberlii in 2018. Another species was described in 2015 based on a supposed crest fragment, but this was later shown to be part of a turtle shell.

<span class="mw-page-title-main">Hațeg Island</span> Prehistoric island

Hațeg Island was a large offshore island in the Tethys Sea which existed during the Late Cretaceous period, probably from the Cenomanian to the Maastrichtian ages. It was situated in an area corresponding to the region around modern-day Hațeg, Hunedoara County, Romania. Maastrichtian fossils of small-sized dinosaurs have been found in the island's rocks. It was formed mainly by tectonic uplift during the early Alpine orogeny, caused by the collision of the African and Eurasian plates towards the end of the Cretaceous. There is no real present-day analog, but overall, the island of Hainan is perhaps closest as regards climate, geology and topography, though still not a particularly good match. The vegetation, for example, was of course entirely distinct from today, as was the fauna.

<i>Hatzegopteryx</i> Genus of large azhdarchid pterosaur from the Late Cretaceous

Hatzegopteryx is a genus of azhdarchid pterosaur found in the late Maastrichtian deposits of the Densuş Ciula Formation, an outcropping in Transylvania, Romania. It is known only from the type species, Hatzegopteryx thambema, named by Buffetaut et al. in 2002 based on parts of the skull and humerus. Additional specimens, including a neck vertebra, were later placed in the genus, representing a range of sizes. The largest of these remains indicate it was among the biggest pterosaurs, with an estimated wingspan of 10 to 12 metres.

<i>Montanazhdarcho</i> Genus of azhdarchoid pterosaur from the Late Cretaceous

Montanazhdarcho is a genus of azhdarchoid pterosaur from the Late Cretaceous period of what is now the state of Montana, United States. Montanazhdarcho is known from only one species, M. minor.

<i>Bakonydraco</i> Genus of tapejarid pterosaur from the Late Cretaceous

Bakonydraco is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Csehbánya Formation of the Bakony Mountains, Iharkút, Veszprém, western Hungary.

<i>Phosphatodraco</i> Late Cretaceous genus of pterosaur

Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.

<i>Aralazhdarcho</i> Genus of azhdarchid pterosaur from the Late Cretaceous

Aralazhdarcho is a genus of azhdarchid pterosaur from the Santonian to the early Campanian stages of the Late Cretaceous period of Bostobe Svita in Kazakhstan. The type and only known species is Aralazhdarcho bostobensis.

<i>Volgadraco</i> Genus of ornithocheiroid pterosaur from the Late Cretaceous

Volgadraco is a genus of pterodactyloid pterosaur from the Upper Cretaceous of European Russia. Volgadraco was originally classified as an azhdarchid. However, recent studies have concluded that it may belong to either the family Nyctosauridae, or the family Pteranodontidae.

Paralatonia is an extinct genus of prehistoric Alytid frog from the late Cretaceous of Sânpetru and Densuş-Ciula Formation, Hațeg Island, modern day Romania.

<i>Alanqa</i> Genus of azhdarchid pterosaur from the Late Cretaceous

Alanqa is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Kem Kem Beds of southeastern Morocco. The name Alanqa comes from the Arabic word العنقاءal-‘Anqā’, for a mythical bird of Arabian culture.

<i>Paludititan</i> Extinct genus of dinosaurs

Paludititan is a genus of titanosaurian sauropod dinosaur which lived in the area of present Romania during the Late Cretaceous. It existed in the island ecosystem known as Hațeg Island.

<i>Gladocephaloideus</i> Genus of ctenochasmatid pterosaur from the Early Cretaceous

Gladocephaloideus is a genus of ctenochasmatid ctenochasmatoid pterosaur from the Early Cretaceous period of what is now western Liaoning, China.

<i>Aerotitan</i> Genus of azhdarchid pterosaur from the Late Cretaceous

Aerotitan is a genus of large azhdarchid pterosaur known from the Late Cretaceous period of what is now the Allen Formation of the Neuquén Basin in northern Patagonia, Argentina.

<i>Eurazhdarcho</i> Genus of azhdarchid pterosaur from the Late Cretaceous

Eurazhdarcho is a genus of azhdarchid pterosaur from the Late Cretaceous period of what is now the Transylvanian Basin of Romania. Its fossil remains dated back 69 million years ago.

Oardasaurus is an extinct genus of lizard from the latest Cretaceous of Romania. It is a member of the Barbatteiidae, a group of lizards closely related to the Teiidae. At 20 centimetres (7.9 in) in length, it was much smaller than the only other named member of the Barbatteiidae, Barbatteius, which lived slightly later. Like Barbatteius, Oardasaurus can be identified by the presence of a crust of bone deposits, or osteoderms, on the roof of its skull; it differs from Barbatteius in the pattern of the sculpturing on this crust. Both Oardasaurus and Barbatteius lived in the isolated island ecosystem of Hațeg Island, having rapidly diversified into various generalist predators of small prey after their arrival on the island during the Early Cretaceous. They went extinct in the Cretaceous–Paleogene extinction event.

<i>Mistralazhdarcho</i> Extinct genus of pterosaur

Mistralazhdarcho is a genus of azhdarchid pterosaur from the Late Cretaceous period of France. The type and only species is Mistralazhdarcho maggii.

<i>Wellnhopterus</i> Genus of azhdarchid pterosaur from the Late Cretaceous

Wellnhopterus is an azhdarchid pterosaur recovered from the Late Cretaceous Javelina Formation in Texas that was previously identified as a thalassodromine. It consists of a set of upper and lower jaws, as well as some cervical vertebrae and a fragmentary long bone. In July 2021, the jaws were given the genus name "Javelinadactylus", with the type and only species as "J. sagebieli"; however, this article has now been retracted. In a paper published in December 2021, the complete holotype was independently named Wellnhopterus, with the only species being W. brevirostris. As of 2022, this is the formal name of this pterosaur.

References

  1. Jipa-Murzea, C., 2012, Upper Cretaceous continental vertebrate assemblages from Metaliferi sedimentary area: systematics, paleoecology and paleobiogeography, PhD Thesis Babeș-Bolyai University, Cluj-Napoca, 228 pp
  2. 1 2 3 4 5 6 Alexandru A. Solomon; Vlad A. Codrea; Márton Venczel; Gerald Grellet-Tinner (2020). "A new species of large-sized pterosaur from the Maastrichtian of Transylvania (Romania)". Cretaceous Research. 110: Article 104316. doi:10.1016/j.cretres.2019.104316.