Aerodactylus Temporal range: Late Jurassic, | |
---|---|
Type specimen | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Aurorazhdarchia |
Genus: | † Aerodactylus Vidovic & Martill, 2014 |
Type species | |
†Pterodactylus scolopaciceps Meyer, 1860 | |
Species | |
| |
Synonyms | |
|
Aerodactylus (meaning "wind finger") is a pterosaur genus containing a single species, Aerodactylus scolopaciceps, previously regarded as a species of Pterodactylus .
The fossil remains of this species have been found only in the Solnhofen limestone of Bavaria, Germany, dated to the late Jurassic Period (early Tithonian), about 150.8–148.5 million years ago. [1] Like all pterosaurs, the wings of Aerodactylus were formed by a skin and muscle membrane stretching from its elongated fourth finger to its hind limbs. It was supported internally by collagen fibres and externally by keratinous ridges. Several well preserved fossils have shown that Aerodactylus was covered in a short, dense coat of bristly pycnofibres, and that it had a rounded triangular crest on its head, as well as a backward-pointing lappet. It is named after the pterosaur-like Pokémon Aerodactyl. [2] The validity of Aerodactylus has been disputed, with some pterosaur experts suggesting that none of the specimens referred to this genus are distinguishable from Pterodactylus. [3]
Aerodactylus is known from six fossil specimens, and though all of them are juveniles, all preserve complete skeletons. [2] The discovery of several specimens with well-preserved soft tissue traces has allowed scientists to faithfully reconstruct the life appearance of Aerodactylus.
The skulls of Aerodactylus were long and narrow with about 64 teeth which were more crowded towards the jaw tips. The teeth extended back from the tips of both jaws, and the tooth row ended before the front of the nasoantorbital fenestra, the largest opening in the skull. [4] Unlike some related species, the skull and upper jaw was curved slightly upward, not straight. [5] A small, hooked beak was present in the very tips of the jaws, with both upper and lower hook no larger than the teeth that surrounded them. [6]
The neck was long, and covered in long, bristle-like pycnofibres. A throat pouch extended from about the middle of the lower jaw to the upper part of the neck. [7]
Aerodactylus, like related pterosaurs, had a crest on its skull composed mainly of soft tissues along the top of the skull. One specimen (MCZ 1505, the counter slab of BSP 1883 XVI 1) shows a roughly triangular soft tissue crest extending upward above the posterior half of the naso-antorbital fenestra and the eye; the crest was 44 to 51 mm long (around 38 to 45% of the total length of the skull) and reached a maximum height of 9,5 mm. [4]
Bennett (2013) noted that other authors claimed that the soft tissue crest of Pterodactylus extended backward behind the skull; Bennett himself, however, didn't find any evidence for the crest extending past the back of the skull. [4] The back of the skull bore a small crest or "lappet" which pointed backward in a cone-shaped structure. The lappet was composed mainly of long, stiffened fibres twisted together in a spiral pattern inside a conical sheath of soft tissue. [7]
The wings were long, and the wing membranes appear to have lacked the furry covering of pycnofibres present in some other pterosaurs (such as Pterorhynchus and Jeholopterus ). The wing membrane extended between the fingers and toes as webbing, and a uropatagium (secondary membrane between the feet and tail) was present, as well as a propatagium (membrane between the wrist and shoulder). [7] Both the finger and toe claws were covered in keratin sheaths that extended and curved into sharp hooks well beyond their bony cores. [6]
In 1850 Hermann von Meyer described the specimen now known by its collection number BSP AS V 29 a/b as a new specimen of Pterodactylus longirostris. Pterodactylus longirostris is a junior synonym of Ornithocephalus antiquus, but Pterodactylus replaced Ornithocephalus through popular use. The specimen BSP AS V 29 a/b was discussed again in Meyer's Fauna der Vorwelt [8] (1860), this time under the name Pterodactylus scolopaciceps. Both Zittel and Wagner took exception to Meyer's new species and it was synonymized with P. kochi in 1883. [9] Broili [10] described a second specimen and used the name P. scolopaciceps, confident that it was a valid species. However, the name slipped into obscurity and Wellnhofer considered it a junior synonym of P. kochi. In 2013 P. kochi was reviewed by Bennet [4] and synonymised with P. antiquus. Vidovic and Martill [2] disagreed with the findings of Bennett [4] and considered the content of P. kochi to be paraphyletic. When Vidovic and Martill separated P. scolopaciceps from P. kochi they considered it so distinct they gave it the genus name Aerodactylus, formed from the Greek words for "wind finger", but chosen in reference to the fictional Pokémon species Aerodactyl, which was based on an amalgamation of different pterosaurian features. [2]
In 2018 pterosaur researcher Christopher Bennett challenged the status of Aerodactylus on several grounds: 1) that the alleged skull features are only trivially different from those seen in Pterodactylus; 2) that the specimens referred to Aerodactylus have an unusual growth regime compared to other Solnhofen pterodactyloids, suggesting that Vidovic and Martill [2] had grouped an unnatural assemblage of specimens; and 3) that no evidence had been presented to rule out effects of preservation or individual variation on the slight differences between Aerodactylus and Pterodactylus [3] . Bennett concluded that Aerodactylus scolopaciceps is a junior synonym of Pterodactylus antiquus. [2]
Pterodactylus is a genus of extinct pterosaurs. It is thought to contain only a single species, Pterodactylus antiquus, which was the first pterosaur to be named and identified as a flying reptile and one of the first prehistoric reptiles to ever be discovered.
Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous. Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger.
Rhamphorhynchus is a genus of long-tailed pterosaurs in the Jurassic period. Less specialized than contemporary, short-tailed pterodactyloid pterosaurs such as Pterodactylus, it had a long tail, stiffened with ligaments, which ended in a characteristic soft-tissue tail vane. The mouth of Rhamphorhynchus housed needle-like teeth, which were angled forward, with a curved, sharp, beak-like tip lacking teeth, indicating a diet mainly of fish; indeed, fish and cephalopod remains are frequently found in Rhamphorhynchus abdominal contents, as well as in their coprolites.
Dorygnathus was a genus of rhamphorhynchid pterosaur that lived in Europe during the Early Jurassic period, when shallow seas flooded much of the continent. It had a short wingspan around 1.5 meters, and a relatively small triangular sternum, which is where its flight muscles attached. Its skull was long and its eye sockets were the largest opening therein. Large curved fangs that "intermeshed" when the jaws closed featured prominently at the front of the snout while smaller, straighter teeth lined the back.
Germanodactylus is a genus of germanodactylid pterodactyloid pterosaur from Upper Jurassic-age rocks of Germany, including the Solnhofen Limestone. Its specimens were long thought to pertain to Pterodactylus. The head crest of Germanodactylus is a distinctive feature.
Ctenochasma is a genus of Late Jurassic ctenochasmatid pterosaur belonging to the suborder Pterodactyloidea. Three species are currently recognized: C. roemeri, C. taqueti, and C. elegans. Their fossilized remains have been found in the Solnhofen Limestone of Bavaria, Germany, the "Purbeck Group" of northeastern Germany, and the Calcaires tâchetés of eastern France.
Campylognathoides is an extinct genus of pterosaur discovered in the Württemberg Lias deposits of Germany; this first specimen however, consisted only of wing fragments. Further better preserved specimens were found in the Holzmaden shale; based on these specimens, Felix Plieninger erected a new genus.
Gnathosaurus is a genus of ctenochasmatid pterosaur containing two species: G. subulatus, named in 1833 from the Solnhofen Limestone of Germany, and G. macrurus, known from the Purbeck Limestone of the UK. Its fossil remains dated back to the Late Jurassic period.
Pterodactyloidea is one of the two traditional suborders of pterosaurs, and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal rhamphorhynchoids by their short tails and long wing metacarpals. The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like Nyctosaurus and Tupandactylus. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles.
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey.
Cycnorhamphus is a genus of gallodactylid ctenochasmatoid pterosaur from the Late Jurassic period of France and Germany, about 152 million years ago. It is synonymous with the genus Gallodactylus.
Diopecephalus is a genus of pterodactyloid pterosaur from the Lower Tithonian of the Lithographic Limestone, Bavaria, Germany. The type and only species is D. kochi, although the name has been applied to Pterodactylus longicollum, with longicollum erroneously listed as the type species.
Altmuehlopterus is a genus of pterosaur belonging to the Pterodactyloidea. It lived in the Late Jurassic of what is now Germany. It was formerly known as "Daitingopterus", a nomen nudum, informally coined in 2004.
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of Pterodactylus and Ctenochasma, and all their descendants.
Aurorazhdarcho is an extinct genus of ctenochasmatoid pterosaur known from the Late Jurassic period of what is now Bavaria, southern Germany.
Bellubrunnus is an extinct genus of rhamphorhynchid pterosaur from the Late Jurassic of southern Germany. It contains a single species, Bellubrunnus rothgaengeri. Bellubrunnus is distinguished from other rhamphorhynchids by its lack of long projections on the vertebrae of the tail, fewer teeth in the jaws, and wingtips that curve forward rather than sweep backward as in other pterosaurs.
Ardeadactylus is an extinct genus of ctenochasmatoid pterosaur known from the Late Jurassic Solnhofen limestone of Bavaria, southern Germany. It contains a single species, Ardeadactylus longicollum, which was originally thought to be a species of Pterodactylus, as P. longicollum.
This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs went extinct millions of years before humans evolved, humans have coexisted with pterosaur fossils for millennia. Before the development of paleontology as a formal science, these remains would have been interpreted through a mythological lens. Myths about thunderbirds told by the Native Americans of the modern Western United States may have been influenced by observations of Pteranodon fossils. These thunderbirds were said to have warred with water monsters, which agrees well with the co-occurrence of Pteranodon and the ancient marine reptiles of the seaway over which it flew.
Cascocauda is an extinct genus of anurognathid pterosaur from the Late–Middle Jurassic Tiaojishan Formation of Hebei Province, China. The genus contains a single species, C. rong, known from a complete skeleton belonging to a juvenile individual preserved with extensive soft-tissues, including wing membranes and a dense covering of pycnofibres. Some of these pycnofibres appear to be branched, resembling the feathers of maniraptoran theropod dinosaurs, and suggesting that pterosaur pycnofibres may be closely related to feathers in dinosaurs.
Propterodactylus is an extinct genus of transitional monofenestratan pterosaurs from the Late Jurassic Painten Formation of Germany. The genus contains a single species, P. frankerlae, known from a complete articulated skeleton. Before its naming, Propterodactylus was referred to as the "Painten pro-pterodactyloid" in the scientific literature.