Monofenestratans | |
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Fossil specimen of Darwinopterus modularis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Clade: | † Pterodactylomorpha |
Clade: | † Monofenestrata Lü et al., 2010 |
Subgroups | |
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Monofenestrata is an unranked group of pterosaurs that includes the family Wukongopteridae and the suborder Pterodactyloidea. [3]
The clade Monofenestrata was in 2009/2010 defined as the group consisting of Pterodactylus and all species sharing with Pterodactylus the synapomorphy of an external nostril confluent with the antorbital fenestra, the major skull opening on the side of the snout. The name is derived from Greek monos, "single", and Latin fenestra, "window". The concept was inspired by the discovery of Darwinopterus , a species combining a pterodactyloid-type skull with a more basal build of the remainder of the body. [3] The Darwinoptera, a primitive subgroup of monofenestratans showing this transitional anatomy, was also named for Darwinopterus and defined as all descendants of its common ancestor with Pterorhynchus . [4]
The earliest known monofenestratan fossils have been found in the Stonesfield Slate formation of the United Kingdom, which dates to the Bathonian stage of the Middle Jurassic, dated to about 166 million years ago. Identified elements include cervical vertebrae, fourth metacarpals and a possible pterodactyloid synsacrum. [5] An earlier undescribed taxon along with the coeval and corregional Allkaruen may also belong to the group, both reported from Middle Toarcian layers of the Cañadón Asfalto Formation of Argentina. [6] [7]
Below is a cladogram showing the results of a phylogenetic analysis presented by Andres, Clark & Xu, 2014. This study found the two traditional groupings of ctenochasmatoids and kin as an early branching group (represented as the group Archaeopterodactyloidea), with all other pterodactyloids grouped into the Eupterodactyloidea. [4]
Pterodactylomorpha |
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Pterodactyloidea is one of the two traditional suborders of pterosaurs, and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal rhamphorhynchoids by their short tails and long wing metacarpals. The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like Nyctosaurus and Tupandactylus. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles.
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey.
Parapsicephalus is a genus of long-tailed rhamphorhynchid pterosaurs from the Lower Jurassic Whitby, Yorkshire, England. It contains a single species, P. purdoni, named initially as a species of the related rhamphorhynchid Scaphognathus in 1888 but moved to its own genus in 1919 on account of a unique combination of characteristics. In particular, the top surface of the skull of Parapsicephalus is convex, which is otherwise only seen in dimorphodontians. This has been the basis of its referral to the Dimorphodontia by some researchers, but it is generally agreed upon that Parapsicephalus probably represents a rhamphorhynchid. Within the Rhamphorhynchidae, Parapsicephalus has been synonymized with the roughly contemporary Dorygnathus; this, however, is not likely given the many differences between the two taxa, including the aforementioned convex top surface of the skull. Parapsicephalus has been tentatively referred to the Rhamphorhynchinae subgrouping of rhamphorhynchids, but it may represent a basal member of the group instead.
Nurhachius is a genus of istiodactylid pterodactyloid pterosaur from the Barremian to Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. Its fossil remains date back about 120 million years ago.
Istiodactylidae is a small family of pterosaurs. This family was named in 2001 after the type genus Istiodactylus was discovered not to be a member of the genus Ornithodesmus.
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods.
Darwinopterus is a genus of pterosaur, discovered in China and named after biologist Charles Darwin. Between 30 and 40 fossil specimens have been identified, all collected from the Tiaojishan Formation, which dates to the middle Jurassic period, 160.89–160.25 Ma ago. The type species, D. modularis, was described in February 2010. D. modularis was the first known pterosaur to display features of both long-tailed (rhamphorhynchoid) and short-tailed (pterodactyloid) pterosaurs, and was described as a transitional fossil between the two groups. Two additional species, D. linglongtaensis and D. robustodens, were described from the same fossil beds in December 2010 and June 2011, respectively.
Rhamphorhynchidae is a group of early pterosaurs named after Rhamphorhynchus, that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley. Members of the group possess no more than 11 pairs of teeth in the rostrum, a deltopectoral crest that is constricted at the base but expanded at the distal end, and a bent phalange on the fifth toe.
Wukongopteridae is a group of basal pterosaurs, found in China and the UK. It contains eight species in five genera, all dated to the Middle to Late Jurassic period,
Archaeoistiodactylus is an extinct genus of wukongopterid pterosaur from the Middle Jurassic of China.
Novialoidea is an extinct clade of macronychopteran pterosaurs that lived from the latest Early Jurassic to the latest Late Cretaceous, their fossils having been found on all continents except Antarctica.
Kryptodrakon is an extinct genus of pterodactyloid pterosaur from the Middle to Late Jurassic with an age of approximately 162.7 million years. It is known from a single type species, Kryptodrakon progenitor. The age of its fossil remains made Kryptodrakon the basalmost and oldest pterodactyloid known to date.
The Mörnsheim Formation is a geologic formation in Germany, near Daiting and Mörnsheim, Bavaria. It preserves fossils dating back to the Jurassic period. It overlies the older Altmühltal Formation in its northwestern extent, and the Painten Formation to the east.
Allkaruen is a genus of "rhamphorhynchoid" pterosaur from the Early Jurassic Cañadon Asfalto Formation in Argentina. It contains a single species, Allkaruen koi.
Liaodactylus is a genus of filter-feeding ctenochasmatid pterosaur from the Jurassic of China. The genus contains one species, L. primus, described by Zhou et al. in 2017. As an adaptation to filter-feeding, Liaodactylus had approximately 150 long, comb-like teeth packed closely together. It is both the earliest known ctenochasmatid and the first filter-feeding pterosaur from the Jurassic Yanliao Biota. Later and more specialized ctenochasmatids differ from Liaodactylus in having longer snouts, smaller openings in the skull, and more teeth. Within the Ctenochasmatidae, Liaodactylus was most closely related to the European Ctenochasma.
Douzhanopterus is an extinct genus of monofenestratan pterosaur from the Late Jurassic of Liaoning, China. It contains a single species, D. zhengi, named by Wang et al. in 2017. In many respects, it represents a transitional form between basal pterosaurs and the more specialized pterodactyloids; for instance, its tail is intermediate in length, still being about twice the length of the femur but relatively shorter compared to that of the more basal Wukongopteridae. Other intermediate traits include the relative lengths of the neck vertebrae and the retention of two, albeit reduced, phalanx bones in the fifth digit of the foot. Phylogenetically, Douzhanopterus is nested between the wukongopterids and Propterodactylus, which is similar to Douzhanopterus in many respects but approaches pterodactyloids more closely elsewhere.
The Cañadón Asfalto Formation is a geological formation which dates to the Toarcian age of the Early Jurassic period of Argentina. The rocks of the formation preserve a diverse biota, including plants, dinosaurs, invertebrates, mammals and pterosaurs, among others. The formation is divided into two members: the lower Las Chacritas Member, and the overlying Puesto Almada member, though the latter has also been assigned to the overlying Cañadón Calcáreo Formation by some authors. The members are typically composed of fluvial-lacustrine deposits consisting of sandstones and shales, with a limestone carbonate evaporitic sequence also being present in the lower of the two.
Propterodactylus is an extinct genus of transitional monofenestratan pterosaurs from the Late Jurassic Painten Formation of Germany. The genus contains a single species, P. frankerlae, known from a complete articulated skeleton. Before its naming, Propterodactylus was referred to as the "Painten pro-pterodactyloid" in the scientific literature.