Thalassodromidae

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Thalassodromids
Temporal range: Early Cretaceous, 112–110  Ma
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DSC 0042.JPG
Replica skeleton of Tupuxuara leonardii
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Clade: Azhdarchoidea
Family: Thalassodromidae
Kellner, 2007
Type species
Thalassodromeus sethi
Kellner & Campos, 2002
Genera
Synonyms
  • Tupuxuaridae
    Martill, Bechly & Heads, 2007

Thalassodromidae or Thalassodrominae (meaning "sea runners", due to previous misconceptions of skimming behavior; they are now thought to be terrestrial predators) is a group of azhdarchoid pterosaurs from the early Cretaceous period of Brazil. They are considered either to be a distinct family within the clade Neoazdarchia, closely related to dsungaripterids or azhdarchids, or alternatively a subfamily within the family Tapejaridae.

Classification

Thalassodromidae traditionally includes only two genera, Thalassodromeus and Tupuxuara , and was defined to include them and any other descendants of their most recent common ancestor. [1] The classification of thalassodromids is controversial. Some studies, including one by Lü and colleagues in 2008, have found that the thalassodromids are more closely related to the azhdarchids than to the tapejarids, [2] and have placed them in their own family (which has sometimes been referred to as Tupuxuaridae, [3] though Thalassodrominae was named first [1] ). Alternately, they have been considered a subfamily (Thalassodrominae) within the Tapejaridae. [4]

Below are three alternate cladograms resulting from studies of azhdarchoid relationships. The first, presented by Felipe Pinheiro and colleagues in 2011, found the thalassodromids (using Thalassodrominae for the group name) as subgroup within the Tapejaridae. The second, presented by Lü and colleagues in 2008, found them to be closer to the Azhdarchidae, though they used the name Tupuxuaridae for the group. The third, presented by Brian Andres, James Clark and Xu Xing in 2014, also found thalassodromids closer to azhdarchids, and as the sister group of the dsungaripterids, however, they used the name Thalassodrominae for their definition. [5]

Azhdarchoidea

Azhdarchidae

Tapejaridae

Thalassodrominae

Chaoyangopterinae

Tapejarinae

Azhdarchoidea

Tapejaridae

Neoazhdarchia

Tupuxuaridae

Chaoyangopteridae

Azhdarchidae

Azhdarchoidea

Tapejaromorpha

Neoazhdarchia

Thalassodrominae

Dsungaripteridae

Chaoyangopteridae

Azhdarchidae

More recently, the Late Cretaceous genera Alanqa and Aerotitan , typically considered azhdarchids, have been recovered as thalassodromids, though their remains are fragmentary enough that this judgement is only tentative. [6]

Related Research Articles

<i>Tapejara wellnhoferi</i> Genus of tapejarid pterosaur from the Cretaceous period

Tapejara is a genus of Brazilian pterosaur from the Cretaceous Period. Tapejara crests consisted of a semicircular crest over the snout, and a bony prong which extended back behind the head.

<i>Tupuxuara</i> Genus of azhdarchoid pterosaur from the Early Cretaceous period

Tupuxuara is a genus of large, crested, and toothless pterodactyloid pterosaur from the Early Cretaceous period of what is now the Romualdo Formation of the Santana Group, Brazil, about 112 million years ago. Tupuxuara is a close relative of Thalassodromeus, and both form a group that is either called Thalassodrominae or Thalassodromidae.

<i>Thalassodromeus</i> Genus of azhdarchoid pterosaur from the Early Cretaceous period

Thalassodromeus is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about 100 million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of Thalassodromeus sethi by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner", and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to T. sethi in 2005, became the basis of the new genus Banguela in 2014, and assigned back to Thalassodromeus as the species T. oberlii in 2018. Another species was named in 2015 based on a supposed crest fragment, but this was later shown to be part of a turtle shell.

Tapejaridae Family of azhdarchoid pterosaurs from the Cretaceous period

Tapejaridae are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain and China, where the most primitive genera are found, indicating that the family has an Asian origin.

<i>Montanazhdarcho</i> Genus of azhdarchoid pterosaur from the Late Cretaceous period

Montanazhdarcho is a genus of azhdarchoid pterosaur from the Late Cretaceous period of what is now the state of Montana, United States. Montanazhdarcho is known from only one species, M. minor.

Gallodactylidae Family of ctenochasmatoid pterosaurs

Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey.

<i>Chaoyangopterus</i> Genus of chaoyangopterid pterosaur from the Early Cretaceous period

Chaoyangopterus is a genus of chaoyangopterid pterodactyloid pterosaur known from a partial skeleton found in Liaoning, China. Chaoyangopterus was found in rocks dating back to the Aptian-age Lower Cretaceous Jiufotang Formation of Dapingfang, Chaoyang.

Eoazhdarcho is a genus of azhdarchoid pterodactyloid pterosaur named in 2005 by Chinese paleontologists Lü Junchang and Ji Qiang. The type and only known species is Eoazhdarcho liaoxiensis. The fossil was found in the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China.

<i>Phosphatodraco</i> Genus of azhdarchid pterosaur from the Late Cretaceous period

Phosphatodraco is a genus of azhdarchid pterodactyloid pterosaur from a late Maastrichtian-age Upper Cretaceous portion of the Oulad Abdoun Phosphatic Basin, Grand Doui, near Khouribga, central Morocco. The type and only known species is Phosphatodraco mauritanicus; the specific name refers to Mauretania.

<i>Huaxiapterus</i> Genus of tapejarid pterosaur from the Early Cretaceous period

Huaxiapterus is a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It is the second genus of tapejarid from this formation, after Sinopterus. Three species are known, though they may not actually form a natural group with each other. It was first named by Lü Junchang and Yuan Chongxi.

<i>Jidapterus</i> Genus of chaoyangopterid pterosaur from the Early Cretaceous period

Jidapterus is a genus of azhdarchoid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. The genus was in 2003 named by Dong Zhiming, Sun Yue-Wu and Wu Shao-Yuan. The type species is Jidapterus edentus. The genus name is derived from Jílín Dàxué or "Jilin University" and a Latinized Greek pteron, "wing". The specific name means "toothless" in Latin.

Azhdarchoidea Superfamily of ornithocheiroid pterosaurs

Azhdarchoidea is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea.

<i>Tupandactylus</i> Genus of tapejarid pterosaur from the Early Cretaceous period

Tupandactylus is a genus of tapejarid pterodactyloid pterosaur from the Early Cretaceous Crato Formation of Brazil. It is notable for its large cranial crest, composed partly of bone and partly of soft tissue. The genus Tupandactylus possibly contains two species, both bearing differently sized/shaped crests that may have been used to signal and display for other Tupandactylus, much as toucans use their bright bills to signal to one another. Tupandactylus crests consisted of a semicircular crest over the snout, and in the case of the type species T. imperator, a bony prong which extended back behind the head. A second species, T. navigans, lacked this prong, and had a much more vertical crest. Soft tissue impressions also show that the small bony crests were extended by a much larger structure made of a keratinous material. The complete crest of T. navigans rose in a sharp, sail-like "dome" high above the rest of the skull.

<i>Shenzhoupterus</i> Genus of chaoyangopterid pterosaur from the Early Cretaceous period

Shenzhoupterus is a genus of chaoyangopterid pterosaur from the Jiufotang Formation of modern-day Liaoning, China. Fossil remains of Shenzhoupterus date back to the Early Cretaceous period, approximately 120 million years ago.

Ctenochasmatidae Family of ctenochasmatoid pterosaurs

Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctively looking teeth, which is thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods.

Chaoyangopteridae Family of azhdarchoid pterosaurs

Chaoyangopteridae is a family of pterosaurs within the larger group Azhdarchoidea.

Germanodactylidae Family of pterodactyloid pterosaurs

Germanodactylidae is a controversial group of pterosaurs within the suborder Pterodactyloidea. It was first named by Yang Zhongjian in 1964, and given a formal phylogenetic definition in 2014 by Brian Andres, James Clark, and Xu Xing. They defined it as the least inclusive clade containing Germanodactylus cristatus and Normannognathus wellnhoferi, which they considered to be close relatives at the time. However, more recent studies by the same researchers have found that these pterosaurs may be only distantly related.

Microtuban is an extinct genus of azhdarchoid pterosaur known from the Late Cretaceous of northern Lebanon.

<i>Vectidraco</i> Genus of azhdarchoid pterosaur from the Early Cretaceous period

Vectidraco, is a genus of azhdarchoid pterosaur from the Lower Cretaceous of England.

Radiodactylus is an extinct genus of non-azhdarchid azhdarchoid pterosaur known from the Early Cretaceous period of what is now Texas, southern United States. It contains a single species, Radiodactylus langstoni.

References

  1. 1 2 Kellner, A.W.A.; Campos, D.A. (2007). "Short note on the ingroup relationships of the Tapejaridae (Pterosauria, Pterodactyloidea". Boletim do Museu Nacional. 75: 1–14.
  2. Lü, J., Unwin, D.M., Xu, L., and Zhang, X. (2008). "A new azhdarchoid pterosaur from the Lower Cretaceous of China and its implications for pterosaur phylogeny and evolution." Naturwissenschaften,
  3. Martill, D.M., Bechly, G., and Heads, S.W. (2007). "Appendix: species list for the Crato Formation." In: Martill, D.M., Bechly, G., and Loveridge, R.F. (eds.), 2007. The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, Cambridge. Pp. 582–607.
  4. Pinheiro, F.L., Fortier, D.C., Schultz, C.L., De Andrade, J.A.F.G. and Bantim, R.A.M. (in press). "New information on Tupandactylus imperator, with comments on the relationships of Tapejaridae (Pterosauria)." Acta Palaeontologica Polonica, in press, available online 03 Jan 2011. doi : 10.4202/app.2010.0057
  5. Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24: 1011–6. doi: 10.1016/j.cub.2014.03.030 . PMID   24768054.
  6. Longrich, N.R., Martill, D.M., and Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS Biology, 16(3): e2001663. doi : 10.1371/journal.pbio.2001663