Nicorhynchus Temporal range: Albian–Cenomanian, ~ | |
---|---|
Holotype jaw fragment of N. capito in multiple views | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Anhangueria |
Family: | † Anhangueridae |
Subfamily: | † Coloborhynchinae |
Genus: | † Nicorhynchus Holgado & Pêgas, 2020 |
Type species | |
†Ornithocheirus capito Seeley, 1870 | |
Species | |
| |
Synonyms | |
Synonyms of N. capito
Synonyms of N. fluviferox
|
Nicorhynchus (meaning "knucker snout", in reference to its likely ecology) is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, N. capito, from the Cambridge Greensand of England, and N. fluviferox from the Kem Kem Group of Morocco. These species were previously assigned to Coloborhynchus . [1]
The genus Coloborhynchus has had a convoluted history, with many species having been assigned to the genus. In 2013, Rodrigues and Kellner considered Coloborhynchus to be monotypic, containing only C. clavirostris, and placed most other species in other genera, or declared them nomina dubia . One of these species was Coloborhynchus capito, which was originally named by Harry Seeley in 1870 as a species of Ornithocheirus . Its holotype is CAMSM B 54625, a snout. In 2001, this species was moved to Coloborhynchus by David Unwin, who also synonymized Ornithocheirus reedi (known from a lost specimen) with it. Rodrigues and Kellner recognized that the species was distinct from Coloborhynchus, but did not give it a new name pending the discovery of more complete material. [2]
In 2018, Jacobs et al. named a new species of Coloborhynchus, C. fluviferox from the Ifezouane Formation of the Kem Kem Group of Morocco based on a partial snout, and also tentatively referred another specimen from the same locality to a different, unnamed species. [3] A 2020 review of a subfamily called Coloborhynchinae by Borja Holgado and Rodrigo Pêgas moved both C. capito and C. fluviferox to a new genus, Nicorhynchus, and also associated the unnamed Ifezouane Formation coloborhynchine to N. fluviferox. The name Nicorhynchus is derived from the Old English nicor (knucker, a kind of water dragon) and the Ancient Greek rhynchos ("snout"), in reference to its likely ecology as a fish-eating, flying reptile found in river and marine deposits. [1]
However, a review of Kem Kem pterosaurs found the traits that distinguish Nicorhynchus from Coloborhynchus to be subtle enough to justify their synonymy, stating that the material was damaged and fragmentary enough to support this. [4]
The species N. capito represents the second largest known anhanguerid (after a Tropeognathus specimen), [5] and indeed the second largest toothed pterosaur known after Tropeognathus . A referred specimen from the Cambridge Greensand of England described in 2011 consists of a very large upper jaw tip which displays the tooth characteristics that distinguish N. capito from other species. The jaw tip is nearly 10 cm (3.9 in) tall and 5.6 cm (2.2 in) wide, with teeth up to 1.3 cm (0.51 in) in base diameter. If the proportions of this specimen were consistent with species of Coloborhynchus, the total skull length could have been up to 75 cm (30 in), leading to an estimated wingspan of 7 m (23 ft). [6]
The describers of the genus, Holgado and Pêgas, had recovered Nicorhynchus within the subfamily Coloborhynchinae, which in turn was within the family Anhangueridae, sister taxon to Uktenadactylus . Their cladogram is shown below. [1]
Anhangueridae |
| |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Cearadactylus is a genus of large anhanguerid pterodactyloid pterosaur from the Romualdo Formation of Brazil, South America. Fossil remains of Cearadactylus dated back to the Albian stage of the Early Cretaceous period, about 112 million years ago. The only known species is C. atrox, described and named in 1985 by Giuseppe Leonardi and Guido Borgomanero. The name refers to the Brazilian state Ceará, and combines this with Greek daktylos, "finger", a reference to the wing finger of pterosaurs. The Latin atrox means "frightful", a reference to the fearsome dentition of the species.
Ornithocheirus is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the United Kingdom and possibly Morocco.
Tropeognathus is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. Tropeognathus is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is Tropeognathus mesembrinus. Fossil remains of Tropeognathus have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil.
Anhanguera is a genus of pterodactyloid pterosaur known from the Early Cretaceous Romualdo Formation of Brazil and the Late Cretaceous Kem Kem Group of Morocco. This pterosaur is closely related to Ornithocheirus, but belongs in the family Anhangueridae. The generic name comes from the Tupi words añanga, meaning "spirit protector of the animals" + wera "bygone".
Siroccopteryx is an extinct genus of anhanguerid pterodactyloid pterosaur, known from middle Cretaceous sediments in modern-day Morocco. Some researchers, such as David M. Unwin, consider the genus a junior synonym of Coloborhynchus.
Coloborhynchus is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. Coloborhynchus is known from the Lower Cretaceous of England, and depending on which species are included, possibly the Albian and Cenomanian ages as well. Coloborhynchus was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related Tropeognathus is now thought to have had a larger wingspan.
Lonchodectes was a genus of lonchodectid pterosaur from several formations dating to the Turonian of England, mostly in the area around Kent. The species belonging to it had been assigned to Ornithocheirus until David Unwin's work of the 1990s and 2000s. Several potential species are known; most are based on scrappy remains, and have gone through several other generic assignments. The genus is part of the complex taxonomy issues surrounding Early Cretaceous pterosaurs from Brazil and England, such as Amblydectes, Anhanguera, Coloborhynchus, and Ornithocheirus.
Lonchodectidae or Lonchodraconidae is a group of pterosaurs within the clade Pterodactyloidea. It has variously been considered to be within Ctenochasmatoidea, Azhdarchoidea and Pteranodontia. They are notable for their high, conical tooth sockets and raised alveolar margins.
Ornithocheiroidea is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods, with fossil remains found all over the world except Antarctica.
Ornithocheiridae is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cretaceous periods, around 140 to 90 million years ago.
The Cambridge Greensand is a geological unit in England whose strata are earliest Cenomanian in age. It lies above the erosive contact between the Gault Formation and the Chalk Group in the vicinity of Cambridgeshire, and technically forms the lowest member bed of the West Melbury Marly Chalk Formation. It is a remanié deposit, containing reworked fossils of late Albian age, including those of dinosaurs and pterosaurs.
Uktenadactylus is a genus of anhanguerid pterodactyloid pterosaurs from the Lower Cretaceous Paw Paw Formation of Texas, United States and the Wessex Formation on the Isle of Wight, England. Fossil remains of Uktenadactylus dated back to the Early Cretaceous period, from about 125 to 100 million years ago.
Ornithocheirae is an extinct clade of pteranodontoid pterosaurs from the Early Cretaceous to the Late Cretaceous of Asia, Europe, North America and South America. It was named by Harry Seeley in 1870 as a family that contains Ornithocheirus and its relatives. The name was emended to Ornithocheiridae, to match the requirements of the ICZN Code that a family-ranked clade should end with an -idae suffix. Brian Andres (2010) in his review of pterosaur phylogeny, defined the name Ornithocheirae phylogenetically, as a node-based taxon consisting of the last common ancestor of Anhanguera and Ornithocheirus and all its descendants. Thus Ornithocheirae is defined to include two families, the Anhangueridae and the Ornithocheiridae, following the opinion of Alexander Kellner and Andres that these families should not be synonymized based on their original phylogenetical definitions. However, subsequent studies in 2019 have found Ornithocheirae to be a more inclusive group containing both Anhangueria and Targaryendraconia.
Lonchodraco is a genus of lonchodraconid pterodactyloid pterosaur from the Late Cretaceous of southern England. The genus includes species that were previously assigned to other genera.
Camposipterus is a genus of pterodactyloid pterosaur from the Early Cretaceous of England. Fossil remains of Camposipterus dated back to the Early Cretaceous, about 112 million years ago.
Anhangueria is a group of pterosaurs belonging to the clade Pteranodontoidea. Fossil remains of this group date back from the Early to Late Cretaceous periods, around 140 to 92.5 million years ago. Anhangueria was named by paleontologists Taissa Rodrigues and Alexander Kellner in a review of Ornithocheirus species in 2013, they defined the clade as a branch-based taxon consisting of all pteranodontoids more closely related to Anhanguera blittersdorffi than to Istiodactylus latidens and Cimoliopterus cuvieri.
Ikrandraco is a genus of lonchodraconid pterodactyloid pterosaur known from Lower Cretaceous rocks in northeastern China and the Cambridge Greensand of the UK. It is notable for its unusual skull, which features a crest on the lower jaw.
Maaradactylus is a genus of anhanguerid pterodactyloid pterosaur known from the Lower Cretaceous period of the Romualdo Formation of northeastern Brazil.
Ornithocheiromorpha is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods, around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing Ornithocheirus, but not Pteranodon.
Aerodraco is a genus of anhanguerid pterosaur from the Albian–Cenomanian-age Cambridge Greensand of England. It contains only one species, Aerodraco sedgwickii. It was originally assigned to the genus Pterodactylus.