Barbosania | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Ornithocheiriformes |
Clade: | † Ornithocheirae |
Clade: | † Targaryendraconia |
Family: | † Targaryendraconidae |
Genus: | † Barbosania Elgin & Frey, 2011 |
Type species | |
† Barbosania gracilirostris Elgin & Frey, 2011 |
Barbosania is an extinct genus of crestless targaryendraconian pterosaur from the Cretaceous Romualdo Formation of the Santana Group of northeastern Brazil, dating to the Aptian to Albian.
Barbosania was named and described by Ross A. Elgin and Eberhard Frey in 2011 and the type species is Barbosania gracilirostris. The generic name honours Professor Miguel Barbosa of the Portuguese Museu de História Natural de Sintra in whose collection the type specimen is present. The specific epithet is derived from Latin rostrum, "snout", and gracilis, "slender", in reference to the slender form of the anterior skull. [1]
The holotype, MNHS/00/85, was originally acquired for Barbosa's personal collection, the basis of the new Sintra museum, from Brazilian fossil poachers. Its provenance is probably the Serra da Mãozinha, [2] implying an Early Cretaceous late Albian age, about a hundred million years old. It consists of an almost complete skeleton including the skull, that is partially articulated and uncompressed. The lower legs and most of the neck are missing. The specimen was prepared in the German Staatliches Museum für Naturkunde Karlsruhe. It is probably of a subadult individual, though the closure of the respective sutures provides somewhat contradictory information, the fusion of some elements indicating it was full-grown. [1]
Several specimens previously referred to Brasileodactylus may instead be individuals of Barbosania. [1]
Barbosania was a medium-sized pterodactyloid with a skull length of 392 millimetres. Its body was 209.5 millimetres long. The skull is elongated with a slight upwards bend in the snout. Its lack of a rostral and dentary median sagittal crest allows a distinction from other related pterosaurs. Also a parietal crest on top of the skull is absent. The authors considered it unlikely that this morphology was age-related and rejected the explanation of such traits by sexual dimorphism unless there was specific evidence to prove this. The describers established a single autapomorphy, unique trait: the possession of thirteen dorsal vertebrae instead of the normal twelve. The first four tooth pairs are extremely long, forming a rosette to catch slippery prey like fish or squids. This rosette is however not laterally expanded, which was reflected in the specific name. There are at least twenty-four teeth in the upper jaw and twenty in the lower jaw for a total of eighty-eight. [1]
Barbosania was assigned to the Ornithocheiroidea sensu Unwin and more precisely to the Ornithocheiridae. [1]
In 2019, Pêgas et al. assigned Barbosania to the clade Targaryendraconia, more specifically to the family Targaryendraconidae, as shown below: [3]
Caulkicephalus is a genus of anhanguerid pterosaur from the Isle of Wight off the coast of England. It lived during the Early Cretaceous period, about 130 million years ago.
Cearadactylus is a genus of large anhanguerid pterodactyloid pterosaur from the Romualdo Formation of Brazil, South America. Fossil remains of Cearadactylus dated back to the Albian stage of the Early Cretaceous period, about 112 million years ago. The only known species is C. atrox, described and named in 1985 by Giuseppe Leonardi and Guido Borgomanero. The name refers to the Brazilian state Ceará, and combines this with Greek daktylos, "finger", a reference to the wing finger of pterosaurs. The Latin atrox means "frightful", a reference to the fearsome dentition of the species.
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Anhanguera is a genus of pterodactyloid pterosaur known from the Early Cretaceous Romualdo Formation of Brazil and the Late Cretaceous Kem Kem Group of Morocco. This pterosaur is closely related to Ornithocheirus, but belongs in the family Anhangueridae. The generic name comes from the Tupi words añanga, meaning "spirit protector of the animals" + wera "bygone".
Tapejaridae are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primitive genera were found in China, indicating that the family has an Asian origin.
Brasileodactylus a genus of pterosaur from the Aptian-age lower Santana formation of Chapada do Araripe, Ceará, Brazil.
Coloborhynchus is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. Coloborhynchus is known from the Lower Cretaceous of England, and depending on which species are included, possibly the Albian and Cenomanian ages as well. Coloborhynchus was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related Tropeognathus is now thought to have had a larger wingspan.
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Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested Tropeognathus.
Unwindia is a genus of pterodactyloid pterosaur from the Early Cretaceous period of what is now modern-day Brazil.
Aussiedraco is a genus of targaryendraconian pterodactyloid pterosaur from the early Cretaceous of Australia.
Guidraco is an extinct genus of toothed pterodactyloid pterosaur known from the Early Cretaceous of Liaoning Province, northeast China. According to many recent studies, Guidraco is a member of the group Anhanguerinae, a subfamily belonging to the larger group Anhangueridae.
Lonchodraco is a genus of lonchodraconid pterodactyloid pterosaur from the Late Cretaceous of southern England. The genus includes species that were previously assigned to other genera.
Banguela is a genus of azhdarchoid pterosaur from the Early Cretaceous period of what is now Brazil. Only one species is known, Banguela oberlii.
Ikrandraco is a genus of lonchodraconid pterodactyloid pterosaur known from the Lower Cretaceous Jiufotang Formation of northeastern China, and the Cambridge Greensand of England. It is notable for its unusual skull, which features a crest on the lower jaw.
Maaradactylus is a genus of anhanguerid pterodactyloid pterosaur known from the Lower Cretaceous period of the Romualdo Formation of northeastern Brazil.
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Iberodactylus is a genus of pterodactyloid pterosaurs belonging to the clade Anhangueria, that during the Early Cretaceous lived in the area of present Spain. The type species is Iberodactylus andreui.