Xericeps Temporal range: | |
---|---|
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Neoazhdarchia |
Genus: | † Xericeps Martill et al., 2018 |
Type species | |
†Xericeps curvirostris Martill et al., 2018 |
Xericeps is a genus of pterosaur from the Late Cretaceous Kem Kem Beds (which date to the late Albian or Cenomanian age) of southeastern Morocco.
The name Xericeps comes from the Ancient Greek : ξερός - meaning dry, referencing the Sahara Desert, in which the pterosaur was first found, and the Latin : cep from capere, meaning "to catch" - alluding to the creature's forceps-like beak. [1]
Xericeps is a medium-sized edentulous (toothless) pterosaur. The term 'medium-sized', in the context of pterosaurs, is generally used to describe pterosaurs with a wingspan of 3–8 metres, and it is likely that Xericeps was nearer the lower end of this range. [1]
The holotype specimen is a partial anterior lower jaw, likely broken off right anterior to where the mandibular rami diverged. The jaw is upturned, with the occluding surface curved in lateral view. On the dorsal surface of the mandibular symphysis are a pair of ridges, similar to those seen in Alanqa and Argentinadraco . [1]
A 2021 study focused on Aerotitan recovers it as a chaoyangopterid closely related to Argentinadraco . [2]
The holotype specimen - FSAC-KK-10700 was discovered by local mine workers at Aferdou N'Chaft, a small mesa near the oasis village of Hassi el Begaa in the Errachidia Province in south eastern Morocco on the Algerian border, and consists only of the pterosaur's fragmented jaws. The specimen was purchased directly at the mine site by British palaeontologist David M. Martill in January 2017, and thus it is possible to confidently establish its precise locality and stratigraphic horizon. [1]
It is believed that Xericeps lived in the mid-Cretaceous period around the Albian-Cenomanian ages (93.9-113.0 Ma). [1]
The holotype specimen's species epithet - "curvirostris" - comes from the Latin curvus, meaning "curved" and rostrum meaning snout, or muzzle. This is due to the specimen's noticeably upward-curved beak. [1]
Azhdarchidae is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well. Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and Titanopteryx. They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms. In several analyses, some taxa such as Navajodactylus, Bakonydraco and Montanazhdarcho were moved from Azhdarchidae to other clades.
Ornithocheirus is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the UK and possibly Morocco.
Thalassodromeus is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of Thalassodromeus sethi by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner", and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to T. sethi in 2005, became the basis of the new genus Banguela in 2014, and assigned back to Thalassodromeus as the species T. oberlii in 2018. Another species was described in 2015 based on a supposed crest fragment, but this was later shown to be part of a turtle shell.
Anhanguera is a genus of pterodactyloid pterosaur known from the Early Cretaceous Romualdo Formation of Brazil and the Late Cretaceous Kem Kem Group of Morocco. This pterosaur is closely related to Ornithocheirus, but belongs in the family Anhangueridae. The generic name comes from the Tupi words añanga, meaning "spirit protector of the animals" + wera "bygone".
Ornithostoma is a genus of pterodactyloid pterosaur that lived during the Early Cretaceous period of Europe, around 110 million years ago. Ornithostoma was once thought to have been a senior synonym of the pteranodontid Pteranodon due to its toothless anatomy and prior naming.
Siroccopteryx is an extinct genus of anhanguerid pterodactyloid pterosaur, known from middle Cretaceous sediments in modern-day Morocco. Some researchers, such as David M. Unwin, consider the genus a junior synonym of Coloborhynchus.
Coloborhynchus is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. Coloborhynchus is known from the Lower Cretaceous of England, and depending on which species are included, possibly the Albian and Cenomanian ages as well. Coloborhynchus was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related Tropeognathus is now thought to have had a larger wingspan.
Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.
The Kem Kem Group is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations, the lower Gara Sbaa Formation and the upper Douira Formation. It is exposed on an escarpment along the Algeria–Morocco border.
Chaoyangopteridae is a family of pterosaurs within the larger group Azhdarchoidea. Chaoyangopterids lived mostly during the Early Cretaceous period, though possible members, Microtuban, Xericeps and Argentinadraco, may extend the fossil range to the Late Cretaceous.
Uktenadactylus is a genus of anhanguerid pterodactyloid pterosaurs from the Lower Cretaceous Paw Paw Formation of Texas, United States and the Wessex Formation on the Isle of Wight, England. Fossil remains of Uktenadactylus dated back to the Early Cretaceous period, from about 125 to 100 million years ago.
Alanqa is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Kem Kem Beds of southeastern Morocco. The name Alanqa comes from the Arabic word العنقاءal-‘Anqā’, for a mythical bird of Arabian culture.
Thalassodrominae or Thalassodromidae is a group of azhdarchoid pterosaurs from the Cretaceous period. Its traditional members come from Brazil, however, other possible members also come from other places, including the United States, Morocco, and Argentina. Thalassodrominae is considered either to be a subfamily within the pterosaur family Tapejaridae, or as a distinct family, Thalassodromidae, within the clade Neoazhdarchia, closely related to dsungaripterids or azhdarchids.
Ornithocheiromorpha is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods, around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing Ornithocheirus, but not Pteranodon.
Aymberedactylus is a genus of tapejarid pterosaur from the Early Cretaceous Crato Formation of Brazil. It contains a single species, A. cearensis.
Keresdrakon is an extinct genus of tapejaromorph pterosaur from the Goio-Erê Formation of Brazil, which dates back to the Late Cretaceous period, 94 to 71 million years ago. Keresdrakon contains a single species, Keresdrakon vilsoni.
Afrotapejara is an extinct genus of tapejarid pterosaur discovered in Morocco. The type species, Afrotapejara zouhri, was named and described in 2020. It was the first tapejarid discovered in Africa and the fourth pterosaur discovered in the Kem Kem Beds.
Apatorhamphus is an extinct genus of azhdarchoid pterosaur from the Kem Kem Group of Morocco. It might have been part of the Chaoyangopteridae. It is only known from a few snout fragments and it likely had a wingspan of between 3–7 metres (9.8–23.0 ft)
Leptostomia is a genus of long-beaked pterosaur from the mid-Cretaceous (?Albian-Cenomanian) of Morocco, North Africa. The type species is L. begaaensis, which was named and described in 2021 from sediments of the Kem Kem Group in Morocco. It was a small animal with a long, slender bill which is thought to have been used to probe sediments for worms and other invertebrates, similar to kiwi birds and curlews. Leptostomia is likely a member of the Azhdarchoidea.
Nicorhynchus is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, N. capito, from the Cambridge Greensand of England, and N. fluviferox from the Kem Kem Group of Morocco. These species were previously assigned to Coloborhynchus.