Xericeps Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Neoazhdarchia |
Genus: | † Xericeps Martill et al., 2018 |
Type species | |
†Xericeps curvirostris Martill et al., 2018 |
Xericeps is a genus of pterosaur from the Cenomanian stage of the Late Cretaceous. It was discovered from the Kem Kem Beds of southeastern Morocco.
The name Xericeps comes from the Ancient Greek : ξερός - meaning dry, referencing the Sahara Desert, in which the pterosaur was first found, and the Latin : cep from capere, meaning "to catch" - alluding to the creature's forceps-like beak. [1]
Xericeps is a medium-sized edentulous (toothless) pterosaur. The term 'medium-sized', in the context of pterosaurs, is generally used to describe pterosaurs with a wingspan of 3–8 metres, and it is likely that Xericeps was nearer the lower end of this range. [1]
The holotype specimen is a partial anterior lower jaw, likely broken off right anterior to where the mandibular rami diverged. The jaw is upturned, with the occluding surface curved in lateral view. On the dorsal surface of the mandibular symphysis are a pair of ridges, similar to those seen in Alanqa and Argentinadraco . [1]
A 2021 study focused on Aerotitan recovers it as a chaoyangopterid closely related to Argentinadraco . [2]
The holotype specimen - FSAC-KK-10700 was discovered by local mine workers at Aferdou N'Chaft, a small mesa near the oasis village of Hassi el Begaa in the Errachidia Province in south eastern Morocco on the Algerian border, and consists only of the pterosaur's fragmented jaws. The specimen was purchased directly at the mine site by British palaeontologist David M. Martill in January 2017, and thus it is possible to confidently establish its precise locality and stratigraphic horizon. [1]
It is believed that Xericeps lived in the mid-Cretaceous period around the Albian-Cenomanian ages (93.9-113.0 Ma). [1]
The holotype specimen's species epithet - "curvirostris" - comes from the Latin curvus, meaning "curved" and rostrum meaning snout, or muzzle. This is due to the specimen's noticeably upward-curved beak. [1]
Azhdarchidae is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well. Azhdarchids are mainly known for including some of the largest flying animals discovered, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and Titanopteryx. They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. Previously it was thought that by the end of the Cretaceous, most pterosaur families except for the Azhdarchidae disappeared from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms. In several analyses, some taxa such as Navajodactylus, Bakonydraco and Montanazhdarcho were moved from Azhdarchidae to other clades.
Ornithocheirus is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the United Kingdom and possibly Morocco.
Thalassodromeus is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of Thalassodromeus sethi by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner", and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to T. sethi in 2005, became the basis of the new genus Banguela in 2015, and assigned back to Thalassodromeus as the species T. oberlii in 2018, though other researchers consider it a valid genus. Another species was described in 2015 based on a supposed crest fragment, but this was later shown to be part of a turtle shell.
Anhanguera is a genus of pterodactyloid pterosaur known from the Early Cretaceous Romualdo Formation of Brazil and the Late Cretaceous Kem Kem Group of Morocco. This pterosaur is closely related to Ornithocheirus, but belongs in the family Anhangueridae. The generic name comes from the Tupi words añanga, meaning "spirit protector of the animals" + wera "bygone".
Siroccopteryx is an extinct genus of anhanguerid pterodactyloid pterosaur which lived in Morocco during the Cenomanian stage of the Late Cretaceous. Some researchers, such as David M. Unwin, consider the genus a junior synonym of Coloborhynchus.
Coloborhynchus is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. Coloborhynchus is known from the Lower Cretaceous of England, and depending on which species are included, possibly the Albian and Cenomanian ages as well. Coloborhynchus was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related Tropeognathus is now thought to have had a larger wingspan.
The Kem Kem Group is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations, with the lower Ifezouane Formation and the upper Aoufous Formation used for the strata on the eastern side of the Atlas Mountains (Tinghir), with the Gara Sbaa Formation and Douira Formation used in the southern Tafilalt region. It is exposed on an escarpment along the Algeria–Morocco border.
Chaoyangopteridae is a family of pterosaurs within the larger group Azhdarchoidea. Chaoyangopterids lived mostly during the Early Cretaceous period, though possible members, Microtuban, Xericeps and Argentinadraco, may extend the fossil range to the Late Cretaceous.
Alanqa is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Kem Kem Beds of southeastern Morocco. The name Alanqa comes from the Arabic word العنقاءal-‘Anqā’, for a mythical bird of Arabian culture.
Thalassodrominae or Thalassodromidae is a group of azhdarchoid pterosaurs from the Cretaceous period. Its traditional members come from Brazil, however, other possible members also come from other places, including Morocco and Argentina. Thalassodrominae is considered either to be a subfamily within the pterosaur family Tapejaridae, or as a distinct family, Thalassodromidae, within the clade Neoazhdarchia, closely related to dsungaripterids or azhdarchids.
Sauroniops is a controversial genus of carnivorous basal carcharodontosaurid theropod dinosaur known from the Late Cretaceous Gara Sbaa Formation, and possibly also the Kem Kem Formation, both of Morocco. The type, and currently only, species is S. platytholus.
Banguela is a genus of azhdarchoid pterosaur from the Early Cretaceous period of what is now Brazil. Only one species is known, Banguela oberlii.
Argentinadraco is an extinct genus of azhdarchoid pterosaur from the Late Cretaceous Portezuelo Formation of Argentina. It contains a single species, A. barrealensis, named in 2017 by Alexander Kellner and Jorge Calvo. Argentinadraco is unusual for bearing a bottom jaw with a concave bottom edge, as well as a pair of ridges and depressions on the top surface. These features distinguish it from all other azhdarchoid groups, complicating its assignment, but it may belong to the Chaoyangopteridae. The ridges on the lower jaw may have been used to feed on small invertebrates in loose sediment within the system of lakes and rivers that it resided in.
Keresdrakon is an extinct genus of azhdarchoid pterosaur from the Goio-Erê Formation of Brazil, which dates back to the Early Cretaceous period, 125 to 100.5 million years ago. Keresdrakon contains a single species, Keresdrakon vilsoni.
Afrotapejara is an extinct genus of tapejarid pterosaur discovered in Morocco. The type species, Afrotapejara zouhri, was named and described in 2020. It was the first tapejarid discovered in Africa and the fourth pterosaur discovered in the Kem Kem Beds.
Apatorhamphus is an extinct genus of azhdarchoid pterosaur from the Kem Kem Group of Morocco. It might have been part of the Chaoyangopteridae. It is only known from a few snout fragments and it likely had a wingspan of between 3–7 metres (9.8–23.0 ft)
Leptostomia is a genus of long-beaked pterosaur from the mid-Cretaceous (Cenomanian) of Morocco, North Africa. The type species is L. begaaensis, which was named and described in 2021 from sediments of the Kem Kem Group in Morocco. It was a small animal with a long, slender bill which is thought to have been used to probe sediments for worms and other invertebrates, similar to kiwi birds and curlews. Leptostomia is likely a member of the Azhdarchoidea.
Nicorhynchus is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, N. capito, from the Cambridge Greensand of England, and N. fluviferox from the Kem Kem Group of Morocco. These species were previously assigned to Coloborhynchus.
Agassizilia is an extinct genus of both freshwater and marine pycnodont fishes from the mid-late Cretaceous. The genus is named after paleontologist Louis Agassiz.
Akharhynchus is an extinct genus of tropeognathine pteranodontoid pterosaurs possibly from the Cretaceous Kem Kem Group of Morocco. The genus contains a single species, A. martilli, known from a small fragment of the premaxillae.